POND PINE WEIGHT EQUATIONS Thesis for the Degree of M. S. MICHIGAN STATE UNIVERSITY FLOYD IL CURTIS 196.9 ’gm‘..._._.,__- LIBRA R Y MIChIf’Q?" I? “c Untvcr r_ “r nW-I m-. THFS'G ABSTRACT This study, conducted in North Florida, presents pond pine (Pinus serotina) tree weight prediction equations for rough green weight to 2-, 4—, and 8-inch top diameters outside bark. The data range is from 4 to 17 inches DBH. Prediction equations are also given for bark weight, peeled green tree weight and the proportion of oven dry wood in the peeled tree. Scatter diagrams with regression lines plotted through the basic data are shown for each of the response variables. Also, the proportion of oven dry wood, at four locations in the tree stem,is shown graphically. Numerous statistics, develOped by multiple regression analysis are included in the Appendix. The prediction equations accounted for 93 to 98 percent of the variation in rough green tree weight and bark weight. The prOportion of oven dry wood variable had less variation explained; at best only 54 percent of the variation in this variable could be accounted for by the dependent variables in the prediction equation. The conclusions of this study are that pond pine tree and bark weights can be reliably predicted within the data range for the study area; that the proportion of oven dry wood decreases rapidly from the butt to the top of the stem; and that the large ii amount of unexplained variation in the proportion of oven dry wood is probably caused by experimental error, natural vari— ation and seasonal variation in moisture content of the tree. iii POND PINE WEIGHT EQUATIONS BY Floyd H. Curtis A THESIS Submitted to Michigan State University in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Department of Forestry 1969 5‘7 /[/22/69 ACKNOWLEDGEMENTS For his encouragement, guidance and counsel in making this study possible, the author is deeply indebted to his major professor, V.J. Rudolph. The author is grateful to The Buckeye Cellulose Corporation of Perry, Florida, for making the data for this study available. For typing and retyping this manuscript, as well as offering constructive suggestions, the author thanks Grace B. Colson. The author's grateful appreciation goes to his wife for being understanding and patient during the preparation of this thesis. iv TABLE OF CONTENTS ABSTRACT . . . . . . . . . . . ACKNOWLEDGEMENTS . . . . . . . TABLE OF CONTENTS . . . . . . LIST OF FIGURES . . . . . . . LIST OF TABLES . . . . . . . . INTRODUCTION . . . . . . . . . POND PINE CHARACTERISTICS . . THE STUDY AREA . . . . . . . . PROCEDURE . . . . . . . . . . DATA ANALYSIS AND RESULTS . . Rough Green Tree Weight . Tree Bark Weight . . . . Proportion Oven Dry Wood DISCUSSION . . . . . . . . . . CONCLUSIONS . . . . . . . . . LITERATURE CITED . . . . . . . APPENDIX . . . . . . . . . . . Definitions of Statistics Presented Multiple Regression Statistics . . Correlation Coefficients Page ii iv vi vii 13 17 24 29 32 33 34 36 42 LIST OF FIGURES Page Figure l. Pond pine study area in north Florida . . . . 5 Figure 2. Data and regression line for pond pine rough green tree weight to a 2—inch top D.O.B. . . . . . . . . . . . . . . . . . . . 14 Figure 3. Data and regression line for pond pine rough green tree weight to a 4-inch top D.O.B. . . . . . . . . . . . . . . . . . . . 15 Figure 4. Data and regression line for pond pine rough green tree weight to a 8—inch top D.O.B. . . . . . . . . . . . . . . . . . . . 16 Figure 5. Data and regression line for pond pine tree bark weight to a 2-inch top D.O.B. . . . 18 Figure 6. Regression line for pond pine peeled green weight to a 2—inch top D.O.B. . . . . . 19 Figure 7. Pond pine bark weight as a percent of rough green tree weight to a 2-inch top D.O.B. by (DBH)2 (Ht.) and DBH class . . . . 20 Figure 8. Average proportion oven dry wood in fresh green pond pine disks by location in the tree . . . . . . . . . . . . . . . . . . . . 23 Figure 9. Data and regression line for pond pine prOportion oven dry wood in peeled green tree 0 O O O O O O O O C O O O O O O O O O O 25 vi LIST OF TABLES Page Table 1. Number of trees in multiple regression program by DBH class . . . . . . . . . . . . 10 Table 2. Mean values of variables in multiple regression program . . . . . . . . . . . . . 12 Table 3. Mean values and standard deviations for sample disks from 61 trees . . . . . . . . . 22 Table 4. Pond pine rough green tree weight to a 2—inch top D.O.B. by DBH and height class . . . . . . . . . . . . . . . . . . . 27 Table 5. Pond pine rough green tree weight to a 4—inch top D.O.B. by DBH and height class . . . . . . . . . . . . . . . . . . . 28 Table 6. Pond pine peeled green tree weight to a 2-inch top D.O.B. by DBH and height class . . . . . . . . . . . . . . . . . . . 30 vii INTRODUCTION This study's purpose is to develop rough green, peeled green and oven dry weight equations for pond pine (Pinus serotina)l/located in the "Big Bend" of north Florida. The basic data were collected in Jefferson, Madison, Lafayette and Taylor Counties during the spring and summer of 1962. Tree stem weight equations, as discussed here, are pre— dicting equations, derived by multiple regression analysis, for estimating merchantable stem weight in pounds. They are similar to predicting equations developed for estimating tree volume in units such as cubic feet, cords or board feet. Tree weights are of interest since considerable quantities of felled timber are bought and sold by weight, especially in the southeastern United States. While this trend toward weight purchases has been primarily with the pulp and paper industry, southern pine sawmills and treating plants are beginning to purchase timber by weight. This trend toward weight and away from volume has occurred because weight measurements are faster, simpler, more consistent and more easily checked than volume measurements. Also, working directly with tree weight eliminates the necessity of using volume-to—weight conversion factors. _L/Scientific and common names are according to Little, (1953). In addition, in the pulp and paper industry, many pro— duction variables such as final product yield, amount of bark, moisture percent, and by-product yield are measured by weight. Weight is of timber also an important factor in regulating the movement over railroads and public highways. Thus, it is logical for those organizations and individuals dealing in timber to This pond pine be interested in tree weight. study presents equations for predicting the following stem weights above the stump: rough green weight to a 2—inch top D.O.B. rough green weight to a 4-inch top D.O.B. rough green weight to a 8—inch top D.O.B.. bark weight to a 2—inch top D.O.B. peeled green weight to a 2-inch top D.O.Bcg/ proportion of peeled green weight, to a 2- inch top D.O.B., that is oven dry wood. While the study is localized, the equations and statistics should be Pond of interest in other areas. POND PINE CHARACTERISTICS pine is one of the minor southern pines. It ranges, in the coastal plain, from the southern tip of New Jersey south- 2 ' = _ .L/By subtraction, yP y2 yB. ward to central Florida and westward to southeastern Alabama. It occurs as far inland as the Piedmont Plateau through the Carolinas and Georgia (Collingwood and Brush, I964). Pond pine is characterized by a comparatively short trunk and a shaggy appearance caused by heavy needle clustering at branch ends. The limbs and trunk have a tendency to produce needle and twig sprouts. Pond pine, unlike most pines, sprouts readily and many stands are established after logging or fire in this manner. The cones are serotinous and consequently cones of varying ages are found on the same tree. The cones Open rapidly after a fire or after the tree is felled. Apparently, fire is necessary for the establishment of well stocked natural stands (Fowells, 1965). Pond pine inhabits the poorly drained mineral and organic soils of the lower coastal plain. On the wet organic soils it often occurs in almost pure stands, while on the poorly drained mineral soils it is found associated with species such as lob- lolly pine (Pinus taeda), slash pine (Pinus elliottii), cypress (Taxodium spp.), and sweetgum (Liguidambar styraciflua). Pond pine growth is slow compared to pines such as loblolly and slash. However, the adverse site on which it occurs often accounts for the poor growth rate. Growth can be improved by the elimination of fire and providing adequate land drainage. THE STUDY AREA The study area is in the lower coastal plain of north Florida (Fig. l). Elevations are generally less than 100 feet above sea level, while the topography is flat and for the most part poorly drained. Lesser vegetation includes wax myrtle (Myrica cerifera), gallberry (Ilex glabra), saw—palmetto (Serenoa repens) and titi (Cyrilla spp.). Pond pine is found primarily on organic soils and poorly drained mineral soils, such asthe Leon and Portsmouth series. The area's climate is typical of the southeastern United States; summers are hot and humid, while the winters are normally dry, with large fluctuations in daily temperature. The mean annual temperature for Perry, Florida, is 68.7OF; the January mean is 55.80F and the August mean is 81.4OF. Annual precipitation for the area is about 55 inches with almost half of it occurring during the months of June, July, August and September. There is an average of 245 days between dates of 32°F or below. PROCEDURE The study was conducted in the spring and summer of 1962 on the lands of The Buckeye Cellulose Corporation. A series of sample points was randomly located on timber type maps. These points were then located in the field and a 4-man crew moved -4- T 11 h ee . i O JaCkSOnville a a ass _ .Perry 0 Tampa Miami . Study Area Fig. l. Pond pine study area in north Florida. 3 Jacksonville Tallahassee o 0 Tampa Miami . Study Area Fig. l. Pond pine study area in north Florida. equipment to the point. The equipment consisted of a small winch equipped crawler tractor, a specially designed skidding and hoisting arch, hydraulic scales with a 3,600—pound capacity, a chain saw, peeling spuds, diameter tapes, numerous small hand tools and miscellaneous materials. Sample trees were selected by making a clockwise sweep with a lO—factor prism and marked by numbering those trees occurring within the sample. Each sample tree had an increment core, reaching to the pith, ex— tracted at breast height (4.5 feet above ground). The increment core was given the same number as the tree, placed in an air— tight bottle, and taken to the laboratory for specific gravity determinations. The sample trees were then felled, keeping stump height as low as possible, limbed and tOpped at 2 inches D.O.B., or where the main stem was no longer identifiable and larger than 2 inches. The following measurements and weighings were then made: 1. Diameter outside bark, at breast height, to the nearest .l-inch. 2. Total tree height to the nearest foot. 3. Bark thickness, at breast height, to the nearest .05- inch. 4. Diameter inside bark, at 16.5 feet above stump, at the butt, at a point one—third of the way up the stem from the butt, at a point two-thirds up the stem from —6— the butt, and at the top of the used stem, all to the nearest .l—inch. 5. Tree age. 6. Rough green tree weight, to a 2—, 4—, and 8—inch top D.O.B., all to the nearest 5 pounds. 7. Peeled green weight of every fifth tree to a 2— inch top D.O.B. The tree stem was weighed on the hydraulic scales by attaching the scales to the tree with two tongs separated by a spreader bar and hoisting the load with the tractor winch line which ran on a fixed roller at the top of the skidding arch. The tree weight was read directly off the scales which had been tared for the log tongs. Every fifth tree was peeled by hand and reweighed. Four 2—inch thick disks were cut from each sample tree at the butt, at a point one—third of the way up the stem from the butt, at a point two—thirds up the stem from the butt, and the top of the used stem. These disks were given the same number as the tree, placed in plastic bags and taken to the laboratory for oven dry weight determinations. The disks cut from each sample tree were brought into the laboratory, loose material was brushed from their surfaces, and they were weighed and recorded fresh green to the nearest .2-gram. -7- Next, they were placed in drying ovens at 1000C. The disk weights were checked periodically until a constant weight was reached, indicating cessation of moisture loss. Most samples reached a constant weight within 32 hours. The proportion of fresh green weight that was oven dry wood was calculated by dividing oven dry weight by fresh green weight. The increment borer used to extract the increment core has a bore diameter of .170 inches which makes the end area of the core .1464 square centimeters. The fresh green volume of the increment core was calculated by multiplying the core length by the core end area. This gave increment core volume in cubic centimeters. The increment cores were placed in drying ovens at a temperature of 1000C. The cores were removed periodically, put in drying chambers and weighed to the nearest .OOl-gram. Weighing had to be accomplished rapidly as the cores quickly picked up moisture from the air. Oven dry weight was reached when increment core weight remained constant. Increment core Specific gravity was calculated by dividing the oven dry weight in grams by the fresh green volume in cubic centimeters. The average increment core specific gravity for 61 samples is .585 with a standard deviation of .059. DATA ANALYSIS AND RESULTS The purpose of this study was to provide pond pine pre— —8— diction equations for the following dependent variables: y2, y4, y8, yB, yP and yOD' There were 57 sample trees used for Y2’ 54 for y4, 29 for y8, 11 for yB, and 61 for YOD' The reasons for using differing numbers of trees is that 61 trees were sampled and used for yOD; some of these trees had data missing and were not useable for y2; 3 trees used in y2 did not have a 4—inch top and thus could not be used for Y47 only 29 trees had an 8-inch top useable for y8; and only 11 trees were peeled for yB. Table 1 shows the number of trees by DBH class used for each dependent variable. After checking the field and laboratory work for errors, the data were punched into cards for multiple regression analysis using a stepwise program on an IBM 7080 computer. Rough Green Tree Weight: The three rough green tree weight response variables, y2, y4 and y8 had eight independent variables used in the program. They were: X1 = DBH, diameter breast height in inches. X2 = Total tree height in feet. X3 = Tree age in years. X4 = Form class or d.i.b. at 16.5 feet above stump. DBH o.b. X = (DBH)2 5 X6 = Logarithm of total tree height. X7 = Logarithm of DBH. Table 1. Number of trees in multiple regression program by DBH class. DBH Variable Class y2 Y4 y8 yB you Inches 4 7 4 9 5 4 4 2 5 6 6 6 6 7 5 5 l 5 8 6 6 1 6 9 6 6 6 2 7 10 4 4 4 l 4 ll 3 3 3 l 4 12 5 5 5 2 5 13 6 6 6 6 l4 1 1 l 0 15 2 2 2 2 16 1 1 1 l 17 1 1 l l 1 Totals 57 54 29 11 '61 -10- X = (DBH)2 (total tree height) = (X )(X ) 8 5 2 The mean values of the variables in the rough green weight programs are shown in Table 2. The program produced predicting equations for those vari- ables with a partial F larger than 1.00. However, for y2 and y4, practically all of the variation was accounted for by the combined variable X8, (DBH)2(total tree height). For ya, in-- dependent variable x5, (DBH)2 accounted for most of the vari- ation. The rough green weight predicting equations and the amount of variation accounted for by the independent variables are as follows: 2 - Response Equation R _§L SyX y 2 Percent Pounds Pounds §2 = .148(X8) - 21 97 57 112 663 4,631 = E8 y4 = .149(x8) - 54 97 54 117 685 4,956 = Q8 ya = 11.38(X5) - 757 93 29 173 878 143.7 ='§5 Appendices 2, 3 and 4 show the multiple regression program output respectively for y2, y4 and y8. These equations are practical in that the transformed values of DBH and height, easily measured characteristics, are used to estimate weight---a value difficult to measure. The inclusion of additional variables only increased the percent of variation accounted for by 1.0, 1.3, and 3 percent, respectively, for Y2' -11- mc0flum>umm90 mo HmQEsz Hm Ha mm gm hm mm¢ mx mmm. I I I I Ammov IAI. Aaxv Iw. was oax mn.a I I I I .um.mw.odH mo Hmooumflomm .le max wow I I I I Ammavxmmov Amxvxaxv max mx mmo. I I I I mmm mo Hmooumflomm 4W. Hax mmm. I I I I hpfl>mum UHMHummm muou .ocH oHumm oax I ms.H I I I mmmcxoanu Mama manned menoaH ax ease emam mass omme Hmoe Aurmammvmxmmao xmeAmxv as I I mmoo.a mama. mmam. mmo mo sssfinmmoq asuaummoq Rx I I moos.a mamo.a osHo.H sesame menu mo eruflummoq ernaummoq ox I I 66.meH em.mm me.mm mxmmov .aH .6m mx _ I I ms. so. me. mmmao shoe oaumm «8 MM me me mm 66 we mmm mane mummw mm _ me I 6 mm we me usage: menu H6869 “was as s.m I m.HH e.m o.m mmo wens mmsucH ax 0mm. I I I I don SoaflIm @003 no GOHuHomonm Oaumm 00» I um I I I do“ gocHIm ufimHQB cmwum omammm mocsom m» I I mum I I mow SUGHIm ufimHQB cemnm Smdom mocsom mm I I omoa 6mm I mow Socfilg unmflws Gmmum Sodom meadow v» I Has HHHH Has moo aou :UGHIN usages cmmum assom weapon as no» ms ms es ms EwuH muHGD mHQmHHm> Emumoum manmfium> mwcommmm .Emumoum COHmmmummH UHQHUHSE as meQMHHm> mo m®5am> new: .N OHQMB y4 and y8. Figures 2, 3, and 4 are scatter diagrams showing the regression lines plotted through the basic data for y2, y4 and y8 respectively. Tree Bark Weight: As only every fifth tree used in the rough green weight analysis was peeled, there are only eleven bark weight sample trees. The independent variables used in the yB program are: X3 = Tree age. X8 = (DBH)2(total tree height) or (X5)(X2). X9 = Double bark thickness in inches. The mean values of these variables are shown in Table 2. As was the case with rough green tree weight, X8 accounted for most of the variation in bark weight. The equation and some statistics are as follows: 2 Response Equation R hi Syx §B X8 Percent -__ Pounds Pounds §B = .o17(x8) + 9 98 11 12 97 5,156 Appendix 5 shows the multiple regression program output for yB. Figure 5 shows the regression line plotted through the basic data. By subtracting the equation for yB from the equation for y , a peeled green weight equation yP is obtained. Thus, yP = 2 _[_—._148(x8) - :I ’Bl7IX8) +_—_9] = .131(x8) - 30. Figure 6 shows -13- Weight in Pounds 2,500 2,000 1,500 1.000 500 Fig. A I y =.ld8£Xq) I—,_l N = 57 Trees r2 = .97 SyIX = 112 Pounds $2 = 663 Pounds 2 = 4631 o// 5)) = 632 ll SX = 4213 l l ' l K I . / ° / /' :K 7/ J .J' 0/ 5,000 10,000 15,000 (DBH)2 (Ht.) = x8 2. Data and regression line for pond pine rough green tree weight to a 2—inch top D.O.B. -14- Weight in Pounds 2,500 2,000 '1,500 1,000 500 Fig. = .149 (x2) - 4 = 54 Trees X = 117 Pounds / = 635 Pounds / xlkfl m r; z I<> M A) I SA = 4137 3. 5,000 10,000 15,000 (DBH)2 (Ht.) = x8 Data and regression line for pond pine rough green tree weight to a 4-inch top D.O.B. -15- Weight in Pounds § = 11.32 (xqi J 7:7 N = 29 r2 = I93 SyX = 173 2,500 Y = 878 SE = 143 . '7 Sy = 648 sx = 54.9 2,000 ,// / / __ // j/l. 1,500 // I O r . 1,000 V/f //’ 1 Z . / 500 . //%/’ . /° // o / 100 150 200 250 (DBH)2 = xs Fig. 4. Data and regression line for pond pine rough green tree weight to an 8—inch top D.O.B. -16- the regression line for this equation. A conservative estimate of Sny is Szyzx + SZyBX = \JI112)2 +~(12)2 = 113. Bark weight as a percent of rough green weight varies con- siderably. Figure 7 shows how this percentage varies over the range of X8. By fitting a curve of DBH over X8 and then super- imposing DBH on X8 in Figure 7, bark weight as a percent of rough green weight is estimated for each DBH class.. Bark percent varies between 19 percent for the 6—inch DBH class and 12 percent for the l6-inch class. Proportion Oven Dry Wood: The oven dry wood in a tree as a proportion of the peeled green weight was obtained by calcul- ation rather than direct measurement, as was the case with rough green tree and bark weight. The calculation was made in the following manner: Let: A1 = Diameter inside bark of the tree's butt disk. A2 = Diameter inside bark of the tree's second disk. A3 = Diameter inside bark of the tree's third disk. A4 = Diameter inside bark of the tree's top disk. D1 = Proportion of oven dry wood in the butt disk. D2 = Proportion of oven dry wood in the second disk. D3 = Proportion of oven dry wood in the third disk. D = Proportion of oven dry wood in the top disk. Since the area of a circle is proportional to its diameter -17- Bark Weight in Pounds § = 017 (x ) ‘ 9 A/ N = 11 [/I rZ = L98 // SyX = 12 [I 250 I = 97 i = 5,156 1/ Sy = 82 // sx = 4,752 // ‘ l 1 200 . /// / 150 /F / ///' 100 l?’ / / o 50 1/ l/. I o 0 5,000 10,000 15,000 (DBH)2 (Ht.) = x8 Fig. 5. Data and regression line for pond pine tree bark weight to a 2-inch D.O.B. -18- Weight in Pounds 9P = 0? -‘9R = .131 (xi) 4 I0 2,500 ~ /// e/ ,/ 2,000 ’//r // [I A/ l , 500 / / 5/ [V’ 1,000 / / / 500 1 z” / 0 / 5,000 10,000 15,000 (DBH)2 (Ht.) = x8 Fig. 6. Regression line for pond pine peeled green weight to a 2—inch top D.O.B. -19- Bark Percent of Rough Green Tree Weight Irk PCrcen = (Qu r §2)(1C0) -25 20 15 \\ \\ \ 10 5 I l l l 1 I l 6 8 1C 12 14 l6 18 I Average DBH . 5,000 101000 15,000 20,000 (DBH)2 (Ht.) Fig. 7. Pond pine bark weight as a percent of rough green tree weight to a 2-inch top D.O.B. by (DBH) (Ht.) and DBH class. -20- squared, the proportion of oven dry wood in the used tree stem, YOD' is estimated by: y = A2 D + A2 D + A2 D + A2 D OD 1 1 2 2 3 3 4 4 A2 + A2 + A2 + A2 l 2 3 4 Mean values for all sample disks are shown in Table 3. The relationships for disk locations within the tree are shown graphically in Figure 8. The independent variables used in the yOD programs are: X1 = DBH in inches. X2 = Total tree height in feet. X3 = Tree age in years. X8 = (DBH)2(tota1 tree height). Xlo = Increment core specific gravity. X11"—L X3 X13 = ___1.-_ x10 X3 A single variable equation using X10, increment core specific gravity, accounted for 45 percent of the variation in proportion of oven dry wood in peeled green pond pine. A three—variable equation increased the amount of variation ex- -21- mmo.H. nos. o.ev mmo.H. mes. N.Hma sooafl sHm. m.mm~ Hoo.H. Hem. o.ssm mEMHO sesumssma 6063 usmsss pumpcmwm mun cm>o cmwuw COHDHOQOHA fimmum QMOE COOS h.mm .m.oma m.¢ma mEMHU sesame sun cw>o Qmmz spasms m.H pews m0 QOB Sumcma Umms m.¢ mouflnuIOBB Spmcma com: v.0 Unflnulmao m.m 005m mGSUCH HmumEMHQ Empm CH Mmfln Goapmooq cmmz MmHQ .momuu Hm Eoum wxmflp mHmEMm How mCOHuMH>mU UHMUcmum Cam mmSHm> cams .m mange -22- Proportion Oven Dry Wood Butt One—third Two-thirds 2-inch used length used length top Disk Location in Tree Fig. 8. Average proportion oven dry wood in fresh green pond pine disks by location in the tree. -23- plained to 54 percent. The proportion oven dry wood equations are: Response Equation R2 EL _§yX ,y 2 Percent l. YOD .642 (X10)+ .151 45 61 .043 .526 .585 2. YOD .489 (X10)+ 54 61 .039 .526 .585 .0025 (X2)- 41.9 .006 (Xl)+ 8.7 .187 Appendices 6 and 7 show the multiple regression program output, respectively, for equations 1 and 2. Figure 9 is the scatter diagram for yOD with the regression line for equation No. 1, above, plotted through the data. DISCUSSION The objectives of the study were partially met with the development of weight equations that explain moSt of the vari- ation in rough green tree weight and bark weight. Unfortunately, the best prediction equation for the proportion of oven dry wood in peeled green pond pine, could, at best, only account for 54 percent of the variation associated with this variable. 2 There may be several reasons for the lack of a high R for yOD' First, for the variables chosen, this may be as much vari- ation as can be accounted for. Second, many detailed measurements -24* .mmnu cmmum beamed CH @003 Sub cm>o coflpuomoum mafia been you mafia coflwmmummn can mama .m .mfim Aoaxv n wufl>mu0 oamaumdm muou ucwEmuocH on. me. om. mm. om. me. 06. o # \\.\ . . . IlY\\\I . . . . . .\\\ Om. . I. .. . ...\ .. . . . . H .\\ . . \\. . . \\.E o \ 4 o IA. . " om. \\ \I . .. mmo. n xm . smo. n »m mam. u m 6mm. u m on 2.0. n Xmm n o N .H 1e m is u z HmH. I Ac.xv m.o. u m -25- pooM K13 uer uorqxodoxd and oven drying could have induced large experimental errors. Certainly, experimental error contributed to unexplained vari- ation. Third, yOD was not a direct measurement, but calculated from four stem samples, assumed to consistently represent the entire tree.' Fourth, there is variation in tree stem moisture content from season to season. Peck (1959) reported seasonal moisture variation on an oven dry basis for loblolly pine, which, when converted to a green weight basis, is as follows: Season Sapwood Heartwood Percent Percent Winter 52.3 37.1 Spring 51.7 32.4 Summer 51.7 36.3 Autumn 50.0 31.9 Bishop and Markworth (1933), also working with loblolly pine, found green weight moisture percent varied between 36.3 percent in May and 42.9 percent in November. The weight prediction equations presented here can be used in forest inventories in the same manner as volume equations are used, or by using Tables 4 and 5, which have pond pine tree weights calculated by DBH class and lO-foot total height inter— vals, respectively, for a 2- and 4—inch top D.O.B. Peeled green weights for a 2-inch top D.O.B. are shown in -26- Table 4. Pond pine rough green tree weight to a 2-inch top D.O.B., by DBH and height class. DBH Total Tree Height in Feet Class 30 40 50 80 Inches -------------------- Pounds —————————————————————— 5 89 26 6 138 191 245 7 196 268 341 8 262 357 451 9 , 338 457 577 10 422 570 718 11 515 694 873 1409 12 ‘ 830 1043 1681 13 978 1227 1977 14 1137 1427 2296 15 1309 ‘1641 2639 16 1492 1870 3005 17 2114 3396 18 2373 3811 A 2 y2 = .148 (DBH) H - 21 N = 57 Trees sampled r2 = .97 Syx = 112 Pounds y = 663 Pounds Mean DBH = 9.0 Inches Mean Height = 43 Feet Sy = 632 Pounds -27- Table 5. Pond pine rough green tree weight to a 4—inch top D.O.B., by DBH and height class. DBH Total Tree Height in Feet Class 30 4O 50 60 70 80 Inches -------------------- Pounds ---------------------- 5 58 I 95 6 107 161 214 7 165 238 311 384 8 232 327 422 517 612 9 308 429 550 671 792 10 393 542 691 840 989 11 487 667 847 1027 1207 12 804 1019 1234 1449 1664 13 953 1205 1457 1709 1961 14 1114 1406 1698 1990 2282 15 1288 1623 1959 2294 2630 16 1473 1854 2236 2618 3000 17 1670 2100 2531 2962 3393 18 2361 2844 3328 3811 A 2 y2 = .149 (DBH) H - 54 N = 54 Trees sampled r2 = . 97 sy,x = 117 Pounds § = 685 Pounds Mean DBH = 9.4 Inches Mean Height = 44 Feet Sy = 628 Pounds -28— Table 6 by DBH class and lO-foot total height intervals. Bark percent by DBH class as shown in Figure 7 may also be used to estimate peeled green weight. To use the yOD equation, an increment core specific gravity sample is necessary. From average increment core specific gravity, an estimate of the proportion of oven dry wood in the peeled green tree stem can be made. In addition to using the equations as mentioned, consider- able statistical information is included. A summary of the multiple regression program outputs are included in the Appendix. In addition, simple correlation coefficient matrices for each problem are in the Appendix, along with definitions (from Steele and Torrie,1960) of the statistics presented. These data should be of value to anyone planning similar studies. CONCLUSIONS The following conclusions are drawn from the results of this study: 1. Pond pine rough green tree weight can be predicted quite reliably for 2-, 4-, and 8—inch top D.O.B. within the data range. The sample trees ranged be- tween 4 and 17 inches. DBH. 2. Bark weight can also be reliably predicted for trees to a 2-inch tOp D.O.B. While the sample size was small, (11 trees), the regression equation accounted —29- Table 6. Pond pine peeled green tree weight to a 2-inch top D.O.B. by DBH and height class. DBH Total Tree Height in Feet Class 30 40 50 60 70 80 Inches -------------------- Pounds ---------------------- 5 68 I 101 6 111 159 I 207 255 7 163 227 291 355 419 8 222 305 388 471 554 9 288 394 394 500 606 10 363 494 625 756 887 11 446 604 762 920 1078 1236 12 725 913 1101 1289 1477 13 856 1077 1298 1519 1740 14 997 .1254 1511 1768 2025 15 1149 1444 1739 2034 2329 16 1311 1647 1983 2319 2655 17 1863 2242 2621 3000 18 2092 2517 2942 3367 A YP = y2 - YB = .131 (Kg) — 30 -30; for 98 percent of the bark weight variation. There is a substantial difference in bark as a percent of rough green weight to a 2-inch top D.O.B. by DBH class. The 6—inch DBH class has about 19 percent of its rough green weight in bark, while the 10— and 17- inch DBH classes have 13 and 12 percent,respective1y. The proportion of oven dry wood weight to peeled green weight is highest in the butt of the tree and lowest in the top. There is almost a straight line relation— ship in the decline of oven dry wood from butt to top. The decline is from .561 in the butt to .407 in the top. The variation in proportion oven dry wood was not satis— factorily explained. Possibly natural variation, ex- perimental errors and seasonal variation in moisture content contributed to the large amount of unexplained variation. -31- LI TE RATURE C ITED Bishop, G.N. and G.D. Marckworth. 1933. Some Factors Influencing Resin Concen— tration in Loblolly and Slash Pines. Jour. Forestry 31(8): 953-960. Collingwood, G.H. and W.D. Brush. 1964. Knowing your Trees. The American Forestry Association. 349 pp. Fowells, H.A. 1965. Silvics of Forest Trees in the United States. U.S.D.A., Forest Service, Agriculture Handbook No. 271. 761 pp. Little, E.L., Jr. 1953. Check List of Native and Naturalized Trees of the United States (Including Alaska). U.S.D.A., Agric. Handbook 41. 472 pp. Peck, E.C. 1959. The Sap or Moisture in Wood. U.S.D.A. Forest Service, Forest Products Labor— atory Publ. No. 768. 5 pp. Steele, R.G-D. and J.H. Torrie. 1960. Principles and Procedures of Statistics. McGraw-Hill Book Company, Inc. 481 pp. -32- APPENDIX -.33-— Appendix 1. Simple regression statisticsl/ y = Predicted value for dependent variable. y = Response or dependent variable. § = Mean of observed dependent variables. x = Independent variable. i = Mean of observed independent variables. N = Number of observations. r2 = Proportion of variation explained by prediction equation = Z(y -§)2 2(y - {NZ Syx = Standard error of estimate = 23y2 - (ny22 = Residual M Sq. EX N — 2 SY = Standard deviation of dependent variable = Eyz - (Zy)2 N N - 1 SX = Standard deviation of independent variable = 2x2 - (EXQZ N N - 1 Confidence limits for a predicted y =‘§ : t Residual MSq)(1 + (x0- x) N Zx31£xf N Where: x0 selected value of x fl II t for given probability level and degrees of freedom for residual mean square. l/For several independent variables see Steele and Torrie, 1960. _34- b coefficient = ny - (§x)(Ey) N 2x2 — (Ex) N 2 Standard error of b coefficient = Residual MS Ex 2 _ (Zx)2 N Confidence interval for B = bl: t Sb Where: t = t for given probability level and degrees of freedom for residual mean square. Standardized = Regression coefficient when each variable b coefficient is in deviations from the mean of its standard deviation. Used for comparing the relative importance of the independent variable = b.§§ 3y _35- Appendix 2. y2 multiple regression program statistics Dependent variable: y2 - rough green pond pine tree weight in pounds to a 2—inch top D.O.B. Independent variable: X8 (DBH)2 (tree height) Number of observations: 57 R2, percent of variation explained: 97 Standard error of y2: 112 Mean of y2: 663 Standard error as a percent of mean: 17 Analysis of variance: Source D.F. Sum Square Mean Square F Total 56 22,397,860 Regression 1 21,707,460 21,707,460 1,729** Residual 55 690,402 12,553 Tabular F .01 with 1/55 df = 7.08 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B coeff Upper Lower Error b Coefficient X8 4632 .148 .155 .141 0004 .984 Constant term in prediction equation = ~21 Equation: y2 = .148 (X8) -21 ~36- Appendix 3. Dependent variable: Independent variable: Number of observations: R2, Standard error of y4: Mean of Y4: Standard error as a percent of mean: Analysis of variance: y4 multiple regression percent of variation explained: CD 54 97 117 684 17 program statistics — rough green pond pine tree weight in pounds to a 4-inch top D.O.B. (DBH)2 (tree height) Source D.F. Sum Sggare Mean Sgpare F Total 53 20,885,770 Regression 1 20,175,480 20,175,480 1,477** Residual 52 710,283 13,659 Tabular F .01 with 1/52 df = 7.19 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B coeff Upper Lower Error b Coefficient X8 4957 .149 .157 .141 .004 .982 Constant term in prediction equation = —54 Equation: Y4 = .149 (X8) -54 .‘-37- Appendix 4. Y8 multiple regression program statistics Dependent variable: Y8 - rough green pond pine tree weight in pounds to a 8-inch top D.O.B. Independent variable: X5 (DBH)2 Number of observations: 29 R2, percent of variation explained 93 Standard error of Y8: 173 Mean of y8: 879 Standard error as a percent of mean: 20 Analysis of variance: Source D.F. Sum Square Mean Square F Total 28 11,752,940 Regression 1 10,941,420 10,941,420 364** Residual 27 811,516 30,056 Tabular F .01 with 1/27 df = 7.68 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B coeff Upper Lower Error b Coefficient X5 143.7 11.38 12.61 10.16 .597 .965 Constant term in prediction equation = -757 Equation: y8 = 11.38 (X5) -757 —38- Appendix 5. yB multiple regression program statistics Dependent variable: yB - pond pine bark weight to a 2-inch top D.O.B.. Independent variable: X8 (DBH)2(Tree Height) Number of observations: 11 R2, percent of variation explained: 98 Standard error of yB: 12 Mean of yB: 97 Standard error as a percent of mean: 13 Analysis of variance: Source D.F. Sum Square Mean Square F Total 10 66,551 Regression 1 65,216 65,216 440** Residual 9 1,335 148 Tabular F .01 with 1/9 df 10. 56 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B coeff Upper X 5157 8 .017 .019 Constant term in prediction equation Equation: YB = .017 (X8) +9 -39-. Lower .015 9 Error b Coefficient .0008 .990 Appendix 6. YOD multiple regression program statistics Dependent variable: yOD - proportion oven dry wood in peeled green pond pine stem. Independent variable: X10 — Increment core specific gravity. Number of observations: 61 R2, percent of variation explained: 45 Standard error of YOD: .043 Mean of yOD: ‘ .526 Standard error as a percent of mean: 8.1 Analysis of variance: Source D.F. Sum Square Mean Square F Total 60 .1947 Regression 1 .0874 .0874 48.07** Residual 59 .1073 .0018 Tabular F .01 with 1/59 df = 7.08 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B Coeff Upper Lower Error B Coefficient Xlo .585 .642 .824 .461 .093 .670 Constant term in prediction equation = .151 Equation: y0D = .642 ()io) +.151 -4o-. Appendix 7. yOD multiple regression program statistics Dependent variable: YOD - Proportion oven dry wood in peeled green pond pine stem. Independent variables: X10 - Increment specific gravity X2 - Total tree height Xl — Tree DBH Number of observations: 61 R2, percent of variation explained: 54 Standard error of YOD: .039 Mean of YOD: .526 Standard error as a percent of mean: 7.5 Analysis of variance: Source D.F. Sum Square Mean Square F Total 60 .1947 Regression 3 .1058 .0353 22.62** Residual 57 .0889 .0015 Tabular F .01 with 3/57 df = 4.13 B coefficients and 95 percent confidence limits: 95 Percent Confidence Limits Standard Standardized Variable Mean B Coeff Upper Lower Error B Coefficient X10 .585 .489 .685 .295 .010 .511 X2 41.9 .0025 .004 .001 .001 .586 X1 807 -0006 -0001 _0011 .003 .366 Constant term in prediction equation = .187 Equation: yOD = .490 (X10) + .003 (X2) — .006 (X1) + .187 _41- ll 2 5m mmo ease mmsa. moma. ms.oe NH. m.ma s.mH m.m soaumssma osmesmum moo «mes mafia. m6-6.H m¢.mm mo. 64 ms o.m sews oo.H mm. mm. ss. so. os. 66. om. 6m. ms oo.H mm. 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