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I. “IN...“ .'.I 7”.M .31.. \‘tllv r $" ."€':'&.. "‘ “Mfg-£75... 125:? i'. --'..- v\ ‘7 .‘ .‘n .. ., . I .1 * ‘ v t -....v N.‘l__ ‘3ijWo.M ‘M’ ‘ I .0- 0 W Zinc! *- 12:81“; biking Em ‘2 5 “i995 ,. «.77 "1-. «a P1 *{fil‘fi'fi'fi “drew-- 'T Mff;“l" :r-vftv-t 1-3 .\:L-:GIC1\¢L bp-{1‘;’J-.L£ JL’ s51 b-vU.-l;_...4-...4 -‘-.a.\J-‘.I.a-A.n .L-LVUT 9.3.1....LJJAL'L ABSTRACT This study was an attempt to relate historical re- cords to ecological factors influencing productivity of ‘HcCoy Creek. Information about the stream's past history was compared to present conditions. The study area was divided into nine units and morphometric data including pool locations and water types were determined for each unit. Where possible, this information was given as the percentage of composition for each unit. Complete data were obtained for the distribution of substrata, and per- manent and non—permanent stream.cover. This was ex- pressed as the percentage of coverage per unit. Physical data were correlated with.biological information obtained from electrofishing, seining, bottom fauna sampling and the relationship between ground water and spawning beds. A questionnaire was used to obtain additional information about the fish population. The main limiting factors appeared to be: a deficiency in available trout spawning facilities, inadequate food production during the critical portion of the year, and lack of protective shelter for adult trout. These deficiencies were compounded by the deleterious effects of man's activities. Recommendations for management are also discussed. AN ECOLOGICAL SURVEY OF A SOUTHERN MICHIGAN TROUT STREAM By Raymond E. Reilly A Thesis Submitted to Michigan State University in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Department of Fisheries and Wildlife 1962 ACKNOWLEDGMENTS I wish to thank Dr. Peter Tack, chairman of the Fisheries and Wildlife Department for his technical advice during the course of this study. Acknowledgments are also due the many local residents who gave freely of their time to volunteer information regarding his- torical sidelights of McCoy Creek. I am indebted to Dr. Gordon Guyer for his assistance in checking the identification of the numerous aquatic insect specimens. Recognition is due Mr. Henry J. Vondett for the in- formation he provided from Conservation Department records on McCoy Creek. I am.also grateful to Mr. Leo Jones, Soil Conservation Service, for his information regarding the McCoy Creek watershed; also the Berrien County Drain Commission and Berrien County Road Com- mission for allowing me to search their records, and the Berrien County Record for supplying the project with publicity. 11 (“'15 :‘ IKTRCDUC‘ Descr Let C11 Get So! Histcfi Int Str Physio Obj Met Ler Hid De; Sub V91 V01 Gra Str Wat Poo Tem Hat BiOIOg Obj Bot1 Dim Bot: TI‘m Pia! E161 Min] GPO] TABLE OF CONTENTS F: m 0 INTRODUCTION. 0 O O O O O O O O O O O O O O O O O O O O O O O O 0 O O O 0 O O O O O O 0 Description of the Study Area.................. Location 0 O O O O O O O O O O O O O O O O O O O O O O O O O O O 0 O O O O O O O Climate.OCOOOOOOOOOOOOOO...OOOOOOOOOOOOOOOO. Geological Description of the Area.......... Soil and Land Description................... History of the Study Area...................... Introduction................................ Stream AlterationSOOOOOOOO.00.0.0000...0.00. \10‘ 0‘ uwwm N H Physical Characteristics....................... 15 Objectives.................................. 15 MathOds...00....OOOOOOOOOOOOOOOOOOOOOOOO...O 15 LengthOOOOOOOOOOOOOOO000......OOOOOOOOOOOOOO 28 WidthOOOO...00......OOOCOOOOOOOOOOOOOOOOO... 28 Deptheeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeee 28 Substrata................................... 31 Velocity.................................... 32 velmeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeee 32 Gradient.................................... 35 stream coverOOOOOCOOOOOOCOOOOOOOOOOOCOOOOOOO 38 water Types................................. #3 POOIBeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeeee hb Temperatures................................ 9 Water Analysis.............................. 5 Biological conditionseeeeeeeeeeeeeeeeeeeeeeeeee 67 ObjectivesOOOOO0.0000000000000000...00...... 67 Bottom Fauna................................ 67 Discussion of Productivity.................. 79 Bottom Fauna and Velocity................... 82 Trent Food Preferences...................... 85 Fishing..................................... 86 Electrofishing.............................. 99 Minnow Seining.............................. 106 Ground water and Spawning................... 11h 111 Sugges Env Tm 83101.33 \ee LIST OF I? LITEPAT’JP. Limiting FactorSOOOOOOOOOOOOOOOOOOOOOOOOOOOOOOO. Suggestions for Management..................... Environmental Improvement................... Trout Populations........................... SMARY. O O O O O O C O C O I O O O 0 O O O O 0 O O O O O O O O 0 O O O O O 0 O O O O O O 0 LIST OF INFORMANTS O C O O O O O O O O O O O O O O O O I O O O O O 0 O O O O O O O LITmTURE CITED. 0 O O O O O O O O O O O O O O O O O O O O O O O O O O O O O O O 0 iv 11. 12. 13. 1h. LIST OF TABLES Geographical features demarcating stream sections.0.00.00...0.0.0...00.000.000.000... Summarization of certain physical charac- teristics of McCoy Creek, by sections....... Velocity of water flow...................... Volume of water flow. By section, in Gable feet/secondOOOOOOOOOOO000.000.00.00... Stream gradient and cross section types..... Percentage of cover based on total stream.1ength and cover..................... ‘Uater types based on total stream length and area.................................... Pool survey indicating their abundance, probable mode of formation, and relative abundance or Shadingeeeeeeeeeeeeeeeeeeeeeeee Horphometric pool data...................... A comparison between water temperature gradients and water shading................. Summarized data for chemical analysis orwaterOCOOOOOOOOOOOOOOOOOOOOOOOO0.0.0.000. Synoptic list of aquatic fauna collected from h9 substrate and h vegetation samples.. Percentage comparisons of the stream's total substrate composition, with samples from.different substratum................... Number and percent of samples by bottom and vegetation types in each food grade..... Page 17 3O 33 3h 39 hl 11.5 h? MB 63 66 7O 77 80 m—s 15. 16. 1?. 18. 19. Her. loca Table 15. 16. 17. 18. 19. Record of trout plantings for McCoy Creek, 19u-8-1962000OOOOOOOOOOOOOOOOOOOOOOO000...... Summarization of catch data from fishermen questionnairesOO...OOOOOOOOOOOOOOOO...O..000 Results of electrofishing by sections....... Minnow seine dataOOOOOOOIOOOO00.0.00...0.... Morphological data for ground water areas locatedOOOOOOOOOO0.000000000000COOOOOO0.0... vi Page 92 97 101 109 118 6T) Firm 1. Ma; boa. Sui 2- KEJ 811‘: sarL 3- Lor )4-12. Dali} nor 13- 1'10:r 11;. Sutr gra 15- Vel tol 16- Re: to 13. 11L. 15. 16. LIST OF FIGURES Map of McCoy Creek showing watershed boundary and relationship to surrounding areaOOOOOOOCOOOOOOOOO000.000.. Map of McCoy Creek showing sectional subdivision locations and various sample site locations..................... Longitudinal profile of McCoy Creek....... Daily water and air temperatures for months of October through June............ Monthly air and water temperatures........ Substrate samples in relation to food gradGSOOOOOOOOOOOOOOOOOOOOOOOOOOOOO.0.0... Velocity and bottom type relationship to insect numbers: by genera............. Relationship of seining productivity to substrateOOOOO00.00000...OOOOOOOOOOOOOO vii Page 19 36 50-58 60 78 8h 112 f.) Plate I Figure igure Plate II Figure Figurg Plate III Figure 1 Fizure 4 Plate IV Figure . Plate I Figure 1. Figure 2. Plate II Figure 3. Figure h. Plate III Figure 5. Figure 6. Plate IV Figure 7. Figure 8. Plate V Figure 9. Figure 10 . Plate VI Figure 11 . Figure 12 . LIST OF PLATES Section I. Natural portion of creek upstream from Curran Road..... Looking east Section II. creek downstream.frmm Curran Road... Section III. from Station BOOOOOOOOOOOOOOOOOOOOOO Section IV. near permanent station.............. Section IV. down Curran Road....... Dredged portion of Looking upstream Looking downstream End of "Bakertown Drain." Downstream from station13......0......0000000000000 Section V. Shows stream emerging from under dense shrubbery.......... One of the more "open" portions or Section VOCOOOOOCOOOOOOOOOOOOO... Section VI. near Station Section VI. near Station Section VII. near Station Section VIII. Looking upstream 17.00.00.00000000000000 Looking downstream 17..................... View'downstream 19.0.000000000000000000 M111 RaCGOOOOOOOOOOOO viii Page 20 2O 21 21 22 22 23 23 25 25 if) Plate VII Figure 1; Figure 1... Plate VIII I Figure 15 Plate IX 1 Fisure 1.: Figure 1' Plate I Figure 14 Plate XI Figure 1: Page Plate VII Figure 13. Section IX. Creek flowing through Clark's property; upstream view...... 26 Figure 1h. Section IX. Upstream.view (dif- ferent location than Figure 13)...... 26 Plate VIII Figure 15. Section IX. Downstream view......... 27 Plate IX Figure 16. McCoy Creek trout catch--19h2........ 89 Figure 17. McCoy Creek trout catch--19h7........ 89 Plate X Figure 18. McCoy Creek trout catch-~195h........ 90 Plate XI Figure 19. Illustrates set-up for obtaining information from fishermen........... 9h ix Much conducted western1 1 only seen. suitabili' cation). Some carried o] I"MAS” C1 cm“); a: "‘1': is ¢ "mu. Aces: cord. do“ ‘3 being I 1879 the 1 They Were chm“ (a; “63‘ in M0) c INTRODUCTION Much trout stream.research and survey work has been conducted in northern Michigan. But there exist in south- western1 Michigan twenty streams, for which, in many cases, only casual surveys have been conducted to evaluate their suitability as trout habitat (Vondett--personal communi- cation). ‘1 Some trout populations and food studies have been carried on for the southern Michigan streams of Augusta and Portage Creeks,‘Kalamasoo County; Donagiac Creek, Cass County; and Silver Creek, Lllegan County. ‘McCoy Creek how- ever, is one of the trout streams receiving only a casual survey. According to Smedley (1938), a survey of early re- cords does not indicate brook trout (Salvelinus fontanalis) as being native to the Lower Peninsula of Michigan. In 1879 the first plantings were made in this peninsula.- they were in southern Michigan; Berrien County was in- cluded (specific locations are not known). in early report (8) states that brook trout occurred in McCoy Creek at least since 1920. Berrien, Cass and Van Buren Counties. i the envir Inn‘s inf present a Itreaa re this the: “MP" l r 2) Survey “Mon, 6 1“Freud W HOCo 8‘1?!” c, “th 11:” “Mush a lad 7 8 803 into the Lj ”Euro 1 I! the "'8. . E Visual evidence of thezmandmade environmental changes are easily noted by any experienced observer. what then is the streamfls present biological and physical condition regarding trout habitat? Can accelerated degradation of the environment be demonstrated? If so, what has been man's influence? If a wide disparity exists between the present and former environment, is it feasible to attempt stream.restoration? These questions form the structure of this thesis. The approach for this study was to: 1) At- tempt a reconstruction of the streamfis past history.' 2) Survey the stream's present physical and biological con- dition, discussing their ecological significance. 3) Pre- pare suggestions, based oniall segments of the study, for improved management of the streamt Description of the Study.Area Location A .UMcCoy'Creek is located in southwestern Michigan's Berrien County. It has headwaters at the Indiana-Michigan state line. The stream.flows in a northeasterly direction through sections 21, 16, 9, h, and 3 in Bertrand Township, and sections 3h, 35, and 25. Buchanan Township; Town 8 and 7, south, respectively; Range 18 west. It empties into the St. Joseph River, which flows into Lake Michigan. Figure 1 represents the streamis relative location in the area. main The inches u average a ring in .‘ uxumm 1 26° (11. s Gsologica Len Hichigan 13 index-1 Period , lost who]. th! lat. Climate The county has a mean annual precipitation of 33 inches,usually evenly distributed throughout the year. The average snowfall is 51.6 inches, more than one-half occur- ring in January and beruary (Kerr g§_gl., 1927). The mean maximum for July is 72° and the mean minimmn for January is 260 (U. S.‘Heather Bureau, 1961). Geological Description of the Area Lane (1907), in his summary of the geology of southern Michigan states that the‘western portion of Berrien County is underlain by Antrim.shale formed during the Devonian period. The surface features of the area were formed al- most wholly by deposition from.the Lake Michigan lobe of the late'Wisconsin glacier, and by shore features of the waters impounded by the melting glaciers. Glacial outwash aprons of gravelly and loam.soils are characteristically distributed between.moraines which have been formed on a north-south pattern. Soil and Land Description A 3 The major—portion of the stream flows through land types characterised by a gently rolling plain dotted with numerous, relatively deep basins which are features of glacial origin. A large percentage of the steeper slopes still remain in forest. Much of the peat and musk retain a cover : cleared ? pasture 1 Il'he tetes G on Pigm- The types we of Ben»: The except be content c The 1‘0 have g 1&3”! a land lhd 1 it Wide 1“ 1 Fox ““110st “Epcot“ 5011 ooh” “d drama lupply or the “my “‘0 ei occurring .; a cover of trees, shrubs and herbaceous vegetation. The cleared land has been used for general farm crops and pasture (Veatch, 193h). The total watershed estimated from a 1927, United States Geological Topographical Survey map and delineated on Figure 1, consists of approximately 15 square miles. The following description of the area's six.main soil types were obtained from.Kerr g§_gl. (1927), Soil Survey of Berrien County. . {A 1 _ The:muck soils are nearly typical in composition except nearer the stream where they may contain a larger content of silt clay or marl than is common. The Plainfield sand and sandy loam.both are reported to have good to excessive underdrainage. The rainfall is largely absorbed and carried away through the substrata of sand and gravel. Drainageways were reported to occur only at wide intervals. Fox loam and sandy loam have unweathered substrate composed of calcareous rocks, gravel, and beds of sand, respectively. The clay in the lemm's B layer results in soil coherence, allowing good but not excessive absorption and drainage. Both soils are capable of retaining a good supply of moisture. Springs are commonly associated with the sandy lcmm. ‘ The survey also mentions Bellefontaine sandy loan, occurring in the steeper slopes, having good underdrainage. Figure 1 Map of McCoy Creek showing watershed boundary and relationship to surrounding area «L ST. JOSEPH RIVER : BucHANA ‘ | l LV— l l I . A I _ l l E I I E l 04; I --.._ -J I Q )l 0' . I I// I / a" [”525 BAKERTWN h -e_ .r «we NILES , 0° ’ 1" I ‘ To new surmise c‘ \ ‘9 RE gr u e“ . a" I Q ‘ to 3 I \ '. °° I o 5 9‘" gouge gasp Q // M HtGAN IND A to I a SCALE 0F Mil—ES 1L» Its what end stony hilly per 3311!. The sting aft This pare generally subsoil a Without a my" Its substrate is composed primarily of unassorted gravelly and stony glacial drift with.mueh limestone. Much of the hilly portions of the study area are composed of these soils. The Miami silt loam.is composed of a silty loam gradu- sting after three feet in depth into heavy unweathered till. This parent till is calcareous but the top and subsoil is generally leached of its line. The drainage is good; the subsoil allowing absorption and distribution of moisture without accumulation of excessive amounts. Histcgy of the Study Area Introduction I Historical data were obtained from.two public agencies and utilizing verbal information from eleven individuals. These individuals have had close contact with the stream through long-tune area residency and/or consistent stream fishing over a period of from five to forty-five years. Their information was compared for accuracy'by cross- checking their data for agreement on certain points. Most informants admitted knowing only about specific sections of the streame Only one person (8) offered information regarding certain conditions of the whole stream, Most queries were directed at obtaining information regarding the altered strewn sections. It appears that these changes had the greatest effect on the overall Q- strean c1 ficent ht The concerniz The Counl the brid; Stream A? Pric Stream cc l lillpo; and an ex BQChanan. aaetion 3 "”1! Is stream character and therefore should be the most signi- ficant historically and biologically. The Berrien County Drain Comission provided data concerning the dates of drainage and cleaning activities. The County Highway Department gave information regarding the bridge and road-work on 11-60. Stream Alterations . ‘Prior'to 1891i most of McCoy Creek followed its natural stream course. The exceptions tothis were the location of [a millpond in section 3 covering approximately 20 acres,2 and an excavated mill race location within the city of Buchanan. It is not known how long prior to 1891; the section 3 millpond existed but it appeared on a deed as early as 18117. In 18911 a group of local residents owning property through which the stream flowed, petitioned the Berrien County Drain Commission to straighten and dredge. the stream comencing from Curran Road (which bisects section 16 in an east-west direction) downstream to the point of Junction of the lew York Central Railroad grade and the Bakertown- Buchanan road. This was a total of 19,011i feet.3 The portion of the stream dredged was to be known as the Baker- town Drain. It was to be maintained by direct tax 2 Atlas Plate Book of Berrien County, 1929. 3 Berrien County Drain Commission Report, 1897. essessmen lowed. first rej benefit f The 1 from dred; (in secti these two in contra 3536 two Buffalo R Stated pr streams r3“ widtf [1111mm ’ 1 12 test me It1 1"" assessment of the landowners through whose land the stream flowed. It is interesting to note that the petition was first rejected because the Commission felt no one would benefit from the dredging. The most marked change in the streamis course resulted from dredging the portion from Curran Road to Buffalo Road (in section 9). The newly straightened section between these two points had a length of approximately 8,000 feet,h in contrast with the original streamis length (between the same two locations) of about lh,000 feet. The work from Buffalo Road to the termination of the drain work, con- sisted primarily of dredging and widening the existing streamfis course. The channel, after dredging, had a sur- face width of approximately 25 feet maximum, to 1h feet minimum, compared to the original surface width of from 12... feet maximum to 5 feet minimum. It is impossible to determine what effect the dredging had on the standing crop and carrying capacity of the stream, since no such records are available. _Data from dredging operations in Indiana (Murray, 1938) .stated that the inediate results were quite striking with a tendency to decrease both the numbers and kinds of organisms present. Bees (1959) in a‘Washington saLmon L h This and all other distances, unless otherwise speci- fied, are based on measurements from an aerial photo map; scale 1:660, obtained with.the use of a map measure. stream at immediate months. E covered 0 titative 1 night hav Prom‘ Hand due 1 in an att. tu1‘31 use and hot m. but“ com. Br 1' “wine t Info ”in! stream study, found a 97 percent reduction in bottom fauna immediately after dredging and a reduced level for 5 months. After 10 months the streamfls bottom fauna had rc- covered completely. lo data is given regarding the quan- titative or qualitative changes in the substrate which might have influenced their overall results. From 189k to 1930 the "drain" remained untouched. Hand dug side ditches were excavated in sections 9 and 16, in an attempt to drain the muck bottom lands for agricul- tural use. The ditches were few in number, small in size and not.maintained (5). However, they undoubtedly contri- buted some siltation to the streamt By 1930 part of the streamfis dredged portion, ac- cording to information from residents (1, 2, 5, 8, 11) again developed many characteristics of its former course. Informants state that upstream.from Buffalo Road, the streamfis substrata and banks were composed of’muck and marl (l, 2, 3, 8). There were many large holes extending hori- zontally into the bank where trout sought refuge (8). The mode of formation of these holes is unknown. 3 I It was reported (1, 2, 8) that the stream bottom from Buffalo Road downstream to the present state highway M-60, consisted mostly of gravel with some sand and silt edges. Several hundred feet of stream, flowing at right angles to the slope of a usually plowed hillside, had relatively more sand than other portions immediately upstream, It {‘75 was sugge. side cont From water cou. formant a of the its structure Versus hi: help acco‘ 61‘1de as ' The gravel mi... 0| 10 was suggested (1) that the frequent erosion of the hill- side contributcd to the deposition of sand. From.the present M-6O to the railroad grade, the water course was described as follows (h, 5, 8, 11). The channel was much deeper and somewhat winding. Each in- formant attempted to verify their conclusion on the basis of the water level then and new related to some physical structure still present in the stream; or water depth versus hip-boot height. This greater reported volume may help account for the deep holes and undercut banks des- cribed as formerly existing. The substrate was composed primarily of medium-sized gravel.mixed with some sand. Little silt occurred along the edges. There were occasional patches of watercress. The creek's channel downstream.from the railroad grade to about the west edge of the city lhnits of Buchanan remained untouched. 3 Approximately 7900 feet (stream measurement from map) east of the west edge of the city, the stream.divides its flow. The original channel flows southeast and the other, called the mill race, flows east-northeast. Both, flowing underground in culverts, again Join under the city of Buchanan and emerge as one from the Portage Road culvert. The mill race was established about 1820 to provide a mill, located in Buchanan, waterpower. The race has been cleaned out by hand and dredged periodically over the years. Prom flows intE taken plad Company's Portage R‘ six feet . level. A‘ is locate feet of 1’; there the dredge?“ th blnkg 9 D8 Ohiy 5118- In 1 undertake 1 bunt as tltiea or the “Me Of Sand e‘ 11 From.the point at which the stream emerges until it flows into the St. Joseph River, only two alterations have taken place; both at an early date on Clark Equipment Company's property. A few hundred feet downstream.from Portage Road, a small pond of about one acre in size and six feet deep had been created. A dam controls its water level. At the east edge of Clark's property, another dam is located creating a cascade with approximately 10-15 feet of fall. Part of the water enters the factory Just above the dam, is used for industrial purposes and then discharged further downstreamt In 1930 the Drain Commission was authorized to re- dredge the stream from the railroad grade upstream to ap- proximately the present M-60 highway. This was a distance of about 5,600 feet. This cleaning removed the undercut banks, natural holes, stumps and logo (11). However, it only slightly modified the width of the stream. In 1932 significant road and bridge construction was undertaken approximately h,600 feet upstream from the Bakertown-Buchanan Road. A heavy duty concrete bridge was built as part of M-60, across McCoy Creek. Large quan- tities of sand and gravel were hauled in to build a roadbed approaching the bridge. This bed is about 15 feet above the surface of the creek and surrounding low land. In order to situate the building forms etc., large quantities of sand and gravel were pumped from the stream.bed; the sand r681 It appeal tron the I result slower eu' In 1‘ creek up sisted o bends th The lentioned Prior to Us: locat This devi race. T'Ho formerly of “tor. haz‘rdoug Van PM}.a1 I'°¢ks 12 sand residue being deposited in and along the stream (ll). It appears that the thick sand deposits in the stream from the bridge to approximately 2,1400 feet downstream are a result of the combined construction activities, and slower current. _ In 1935 additional work was done by the VPA on the creek'upstream from 14-60 to Buffalo Road. This—work con- sisted of removing logs, cutting brush and removing stream bends that had reformed in the creek (1, ll, 9). The mill race was dredged by dragline in 19h1. As mentioned, this race had been hand-cleaned many times, prior to 1911.1. A water diversion structure (now defunct) was located where the mill race and main channel separate. This device regulated the amount of water entering (the mill mo. ‘i'wo observers (8, 11) noted the main branch was formerly (prior to dredging) a very fast flowing stretch of water.“ Due to its velocity and depth, wading was hazardous. .Undercut banks were common. The bottom type was primarily medium to large gravel, scattered with large rocks. The water area was heavily shaded by long and dense overhanging sedge. The bottom is now primarily sand. The undercut banks are gone. The swiftness and volume of water has been greatly reduced. Whether the dredging of the mill race and abandonment offlthe water diversion structure were entirely water is approxime dredged the etre driven b 13 entirely responsible for the reported decreased flow of water is speculative. In 1936 the Pike Lake outlet flowing into the creek approxbmately 600 feet dcwnstream.from Buffalo Road, was dredged and straightened. The original outlet flowed into the stream.and must have carried a fair volume of water driven by a velocity great enough to scour a good sized pool at the point of Junction with the creek. This pool was used as a swimming location (1). The bottom.was firm and the water approximately h feet deep and cold. Almost immediately after this outlet was dredged, the pool became unsuitable for swimming due to the deposition of muck and silt dislodged during the dredging. The wider outlet, now having less velocity, apparently could not keep the pool scoured out. According to the Drain Commission's records, scattered work was done on the stream.during l9hl and l9h2. It is not known where this work was carried out. ‘ A small one-quarter acre pond was dug within the last 15 years, 3,200 feet westerly from the mill race sep- aration. It is fed by a diversion channel leading from the creek into the pond. The outlet goes back into the creek a short way down from.the pond's inlet. Only minor scattered cleaning work has occurred on the main stream since l9u2. Appr and a an? upstream the land: extremely below the data to (3 original recent or In 1 We]! land attempt t b! dredg: Channel. approxima Frog”! ‘1 in a 31m Acco section 0 110110",e d ailted in 8° heavu in Many a 1h Approximately 11 years ago, a portion of the strewn and a small pond was dredged out about one-quarter mile upstreamifrom the Bakertcwn-Buchanan Road. According to the landowner (7), the dragline operator commented on the extremely thick layer of gravel occurring about 3 feet below the present muck substrate. There is insufficient data to definitely say that this gravel was the streamis original substrate and the 3 feet of muck is a result of recent erosional deposition. In 1955 an agricultural venture was initiated on the muck lands between Curran and Buffalo Roads. This was an attempt to drain the land for use in a corn growing program by dredging a network of side ditches at angles to the main channel. The work between 1955 and 1960 accounted for approximately 5 lineal miles of ditches being dug. This program.was a failure and abandoned by 1960. Some interest in a similar project has been revived by new landowners. According to a local resident (8) who has fished this section of the stream.for about 30 years, all the former hollowed out holes extending horizontally into the banks silted in. This area's bottom,banks, and pools are now so heavily silted over that navigating the stream by canoe, in.many stretches, is very difficult or impossible. Its character is now that of a ditch.more than a stream. A previou. for McCc analysis The length, the 8008 Signific of large water cr< bank cove debrig, ‘ ersce dolly se The six-gal ' or these Mm tr 8amPlea a w . The o°t°ber t; continued i r 15 Physical Characteristics Objectives A survey of the published literature revealed no previous work of an intensive nature having been completed for McCoy Creek. A prime objective then was a thorough analysis of the streamis physical characteristics. The stream survey included: 1) ascertaining its length, average width and depth. 2) Sketching in detail the geographical course of the stream. 3) Mapping in all significant characteristics such as location and abundance of large beds of aquatic vegetation, and in the case of water cress, its water surface area coverage; location of bank cover and its degree of stream.ehading; logs, stream debris, undercut banks; pools-~their approximate water surface area and depth, and types of substrate. 3) Ran- domly selecting stations throughout the stream's length. The data to be collected at each station included: stream velocity, width, depth, and substrate types. Some of these stations were to be later used as collection. points to obtain water for chemical analysis, bottom.fauna samples and minnow samples. Hethods The major portion of this survey extended from mid- October to the first of November, 1961. Certain phases continued into the summer of 1962. A. c I depth taken da Air and afterno: taken fr inches b will be peratura l’or' dISSeI-ta into see This and mess: Iine gec1 graphics] and the d Their re Ille length e Buffalo F' Where he the “tree Plant to In the“ p0013. 16 A convenient permanent station site was selected and a depth gauge installed in mid-stream. Readings were taken daily; the water depth recorded in tenths of a foot. Air and water temperature were recorded morning and late afternoon at the same location. ‘Hater temperatures were taken from the bank, with the thermometer held several inches below the surface of the water. Further details will be given and discussed under the topic of "temp peratures." . For convenience of reference during the course of this dissertation, a discussion of the division of the stream into sections will be given at this point rather than later. The stream.was divided into sections based on visual and measured differences noted in the streamfls character. line sections were defined. Table 1 gives their geo- graphical points of demarcation, the length of each section and the dominant characteristics used to identify'theme Their relationship to the stream.1s shown in Figure 2. The survey was accomplished by wading the streamis length except for the following areas: upstream.from the Buffalo Road bridge to the headwaters. A canoe was used where practical, and the remainder was surveyed by walking the streamis bank. Downstream from the Buchanan sewage plant to the St. Joseph reservoir the bank was traversed. In these areas stream depth was recorded only in larger pools. Table 1 Geographical features demarcating stream sections Innrvnflmnifl ..__“._Lcn- nhnrncteristlcfi 17 . noon nacho nope: ammonnu tonne: van Hosanna ova: .wuamene hops: ho pnzoad suave: somewhat nausea Sonioeom Em .Eeonpn wnacnus_aeaawano on emmahn mecca: 0005 o H> embahn neooos n.3eaun< song Hennsmo seamen .newme aaam Buchanan pooh .wnumenu memo: no masons emaoq com on oeom use» .Eoenpu wuaoaat Headwano inexemunememusm come w > . -upneHm cassava no Aeoenw neon npzonw canon .noaac emanaeno laden new boom eeHnEeuom .oHoom renunsoau mendnoamnnzop unmaeaam wwnficema never eduuaa inexem on 00:! coed. .m >H . eensoo ranches Housman e eeansenem 00:: on .mnucemu some: no masons shame: oeom caehnsm com: .d HHH -ne>oc ounce and : ewcunn ocean e333 .sooeon Eon: eeom Sudden .nopav emenaehn seanaenem on poem damage 000» N HH enema Henupea ma hannem 41 neom_nenaso o» wnaonaslahnnsnninsoahen knob naeuesueen.aonm ooao H H . noun-endaec no Hpeeh mam .ow noaananepoeAeme unannaswsaueun amass decanneamoom damned casuah noaaeem , A . y . w A . emmaauonmnw . J II-Juoztiooeoc emLoE Hmouzaoawoeo Ideal-1141!!! I . nomcofl, tandem soeoeon cu IEHNOLQQ‘ 18 moon» omnoa no hmonee hp mnaconm .uuoammnw macaw .uoupsaaoa owusou no\pnm assumescna on neonnse noon non nouaooe no pros haaeoanhnm nsoonomoaon cop non swoonpam nebam smooch .am on canard emeaaom 0000 MA NH season ream .muaoeno none: onooen .Heuee seamen neanaeuom owne>< stennano on eoom Haas ooom NH HHH> museums nosed serene» send endow .noeae seamen .nOaphom memos ma madman» hope: no unsoaw sauces --wueoue3 as._seonee Handmade canard when on mesons can: com: , HH HH> neapnaaopeenano msunaasmsaauan moancenaaev no exams Heoanmeawoeo Auto.“ :3 newton epmmmwonamd .en enamah qoaaoem ,Figure 2 Map of McCoy Creek showing sectional subdivision locations and various sample site locations '3‘, \q . ‘ e - section ‘\ “‘ $ECI'ION Buchm- '11 $1“ \f'mT Section ‘21 ‘ \ ‘\‘ ‘m \ . _ M 56010:; ---~- ..5 52m? '9 . e . Sac-noel I . .1 5 ‘/ SGCTTON 1'! SC!“ on. GO Sec-now m: / .9!" 3f ('1 L9“ . SGC‘UW n 5 we. (I E LEGEND S‘suaemm Focus; Somme L--- J? E~ {\ectaoeish'we semen m- (“Ransom SG‘Na fihmefle r. Mann-my TQM peRnWRé Pa Pcamaueur 'SYRTRON 5““‘W I ---'- section BOUNDRRY Q i j.- Sceic o? maes a 7"? I'm _ hrs-1 Iilit'h'llr'e Fleur 20 % Plate I '3‘. {"3 ' ’1'.' 14‘s Figure 1. Section I. Natural portion of creek _. upstream.from Curran Road. Figure 2. Curran Road looking east. Bump in road indicates presence of underground springs. .1 l“ Figure - P181135 ’4 A 21 Plate II .Pigure 3. Section II. Dredged portion of creek downstream.from Curran Road. 896! NV! {P J f fv J‘fifv wiry—Ky" \‘ Figure h. Section III. Upstream view near Station 8.. '1' 1Euro 5 ' “an. 6 Figure 6 . L 22 Plate III 8961 NY! 1 ‘ \ Section IV. Looking downstream towards permanent station. 8961 NY! " "— -\' “- Section-XIII.- End "of "Bak‘ertown Drain." Downstream view fromStation 13. ,- I“ Figure 7. Figure 8 23 ‘ Plate IV a", 8961 NV! \ .Pigure 7. Section V. Shows stream.emerging from . under dense shrubbery. 8961 NY! } Pigure 8. Section V. One of the more "open” 8 . portions of this section. a , ' ”We 10, Mem 9. Se new 2h , Plate V 'V 8961 NV! '0' .34 _ ’ . '.A.““ ( I I .l ,' ‘ ‘; ‘:-‘~4_.;‘”‘;'; 115‘. ‘1. air { ‘ '1‘“ fi 4». .‘ FN--‘-\ V a to. ‘ I . J ' ' 'A; ’ its ’ h f' 5 r ’ " '4 ' 15; W\‘ ,‘i a -7) ‘8 3" , Figure 9. Section VI. Looking upstream toward . Station 17. 896! NY! Figure 10. Section VI. Looking downstream.near .. Station 17. Note dense mate of water cress. Dotted lines indicate approxi-g mate extent of ice coverage. § '9' ”Sure .- Figure 11. .Figure 12. 25 Plate VI 9961 KY! ——\K ’ ‘kln Section VII. View downstream near Station 19. Dotted lines indicate approximate extent of ice coverage. 2296! XV! \Mill Recon-Section VIII: This section completely frozen over during part of winter. Figure 13 Fl1811M 1h 0"" 26 .Plstc VII 896! NY! Figure 13. Section II. Upstream tiew through . Clark's property. 9961 NY! f" ' Ward w ' o sir-V1.5? '\ R‘k" er?- 35" ‘Ws‘ Figure 11;. Section IX. Upstream view) through . Clark's property. 27 Plate VIII £961 NVl’ Figure 15. Section IX. Downstream view through Clark's property. Dotted lines indi- cate approximate extent of ice coverage. 28 A description of the methods used to measure the streamfis physical characteristics, and discussions of each, follows. Length " See footnote h, page 8. use This measurement was made at the selected stations only from solid bank to bank. Readings were recorded to the nearest one-half foot. Beds of water cress were in- cluded as part of the strewn since they were growing in the water. A total of 27 measurements were taken. The average widths per section is summarized in Table 2.- Results and Dieoueeion.--widths in Sections II, III, and IV show the results of earlier dredgings. They are much greater than would normally be expected compared to relative widths in adjoining upstream.and downstream sections. It appears that this stream width change has decreased the streamis velocity from that which.wou1d have occurred had it not been widened. 222m ‘Ilneasurements were taken frequently while wading the stream in: a) the main channel and/or the center of the stream, b) all pools. c) 8‘ The deptl center a; All 1 nearest 1 clear waf In those the sele< figsgg Table 2 e V compare that the Stream 13. greater 1 Itcent 8e Field depth “I The chanr much or 1 ”Minder "”880 d Present P be 8118330 5\ EdSe d 6 The at “Dy u: 29 e) stream edges5 The depth was also recorded at each station for the stream center and at each edge. All measurements were made with a yardstick, to the nearest inch. In the case of stream edges, only the clear water flowing over the semi-solid silt was measured. In those sections not waded, depth was recorded only at the selected stations. ‘Results and Discussion.--Analysis of the data in Table 2 shows a noticeable difference in depth in section V compared to other adjoining sections. The data suggests that the bank cover has acted as a stabilizing agent. The stream.basin in this section is therefore subject to greater scouring action (in the main channel) than ad- jacent sections. ‘ Field data show this data section to have the greatest depth variation between the main channel and the edges. The channel was approximately 31 to 3k inches throughout much of its 1ength.but usually only 3-h feet wide. The remainder was quite heavily silted bringing the depth average down considerably. How much of this silt was present prior to "recent" dredging activities can only be guessed.6 ‘ 5 Edge defined here to mean the approximate mid-point between center and bank. 6 The stream.was randomly divided into units; one to many units composed a section. 'Within each.unit Table 2 Summarization of certain physical characteristics of McCoy Creek, by section on fifggompggjfion averageffiecf substrata-.erce 30 m 0 ma : mm mm ooo.om _~upma mm m mm .. mm 0H Hm m.~H coco le -- -- mm -- on m: ma m.:a coon HHH> ma ma 0: a -- mm Ha 0.:H com: HH> s as mm o :H mm aH m.aH coma H» a 9 mm a a me am o.MH come > -- : ma ma 3: ma :H o.ma coo: >H. -- e m -- me am mH o.mH com: HHH -- -- m -- :H as 0H o.o~ coop HH -w -4 u» -w om ow m m.m ooao H mxmwh Hobmrm HobquWrndh cmmw aHm» ammw umumuw “MM“: nwmmwa uncuuoom HHmmm summon was waam tunam .‘ owaAo>< emanobd noaaoom nOHvoom\omaho>a nomwmnomaooAmmoOhomrncnuhumnvm Subst deter] by ac: were: the sa silt 11 but son Proport with an sand 1m Due sewage I three pg Tab tota1 d1. A mltum ‘ Sand-81 percent 8 31 Substrate Results and Discussion.--The bottom character was determined visually and by the degree of abrasiveness felt by securing the bottom with the foot. The types mapped were: silt; sand and silt mixture (usually stratified with the sand overlaying the silt); sand--usually some mineral silt intermingled; sand--gravel.mixture--usually fine gravel but some medium-sized gravel intermingled--always a large proportion of sand; medium-sized gravel-~mostly occurring with an admixture of sand; small rocks--normally little sand intermixed. Due to the turbidity of the water below the Buchanan sewage plant, the bottom types were not mapped except at three points. Table 2 shows that silt composes 33 Percent of the total different types of substrata (25 percent loose silt7). A mixture of sand-silt makes up 32 percent, sand h percent, a sand-gravel mixture 18 percent, mediumrsized gravel 8 percent and small rocks 5 percent. certain physical characteristics (e.g., water types, water cross beds, etc.) were mapped as to their approximate area of water coverage. This was based on their visually esti- mated width from the bank towards the streamis center, times the lineal distance they occurred in a streamis unit. Areas of physical characteristics per unit were combined to compute their percent occurrence for a total stream.section. 7Loose silt refers to silt not forming soil media for watercress beds. Velocit‘ Cu: using a of nylon extend t] A series The surfs time in 5 Table 3 8! R_e_s_u_g moderately average ye for Sectio. 8fictions (1 III) "”38 checked, M on “19 Hate. cause, s cm do" the cez % This 33 (19510 metho‘ was collect“. linal Point O 32 Velocity Current was measured in midstream at 27 stations using a light plastic float attached to a 5 foot length of nylon line, .01 in diameter. The length of time to extend the line's full 5 feet was measured by stopwatch. A series of 3 trials were made and the results averaged. The surface velocity was computed by dividing the average thme in seconds into the distance traversed by the float. Table 3 summarizes the data for the different sections. Results and Discussion.--On1y two sections had a moderately swift (approximately l-l.5 feet per second) average velocity. A two foot per second average velocity for Section IX could be considered swift water. Three sections (I, III, IV) were moderately slow and two (II and III) averaged slow water. The headwater section was not checked. Much of the dense shrub cover in Section V lies on the water's surface along the stream's sides. This causes a current speed-up by channeling most of the water down the center of the strewn. Volume (This was determined, using the Robins and Crawford (l95h) method, as outlined by Lagler (1956). This data was collected at 7 locations near the geographical ter- minal point of the sections designated in Table h. Table 3 Velocity of water flow Flow CPS Less th; .S-.9 1- 1.1L 1.5 - 1. 2+ 33 . __§ections Flow in I I I I_l_V .. IX . CFS Averagg_velocity:;feet[secondi_ Less'than .5 x ' ' x ,5 - .9 x x x l - l.h X X 105 - 1.9 X 2+ X Table h Volume of water flow By section, in cubic feet/second cps—r N01 measx 34 *Section CPS I Not Not measured measured 3.3 5.0 7.3 11.0 6.0 nth 17.0 D“ Gradient A t contour ' gradient of strea: contour : and the '. R_e_e_1 the stre: the gradi course 1, one thou: increase; Streamva thousand‘ Alle that in E Der one t GI“filing feet Per do"intuit 1.5 feet one thong A11s that “1th (mud to 35 Gradient - A topographical map (U. 8. Geological Survey) with a contour interval of 20 feet was used to determine the gradient of the stream. It should be noted that the rate of stream.fall is not necessarily uniform between mapped contour intervals. This is especially true for Section VII and the lower part of Section VI. Results and Discussion.--A longitudinal section of the stream.1s pictured in Figure 3. This illustrates that the gradient for the upperBh percent of the streamls course is quite uniform, averaging 1.2 feet of fallper one thousand feet of stream length. The average gradient increases sharply for the remaining 16 percent of the streamis course; the gradient being 5.9 feet per one thousand. Allen's (1951) reference to Huet's (l9h6) work stated that in Belgium, waters with a gradient greater than 3 feet per one thousand are predominantly occupied by trout. Grayling and chub also occur in gradients less than 6.5 feet per one thousand. He further mentions trout being a dominant species in streams having gradients as low as 1.5 feet per one thousand. Between this and a .75 feet per one thousand trout occur in only small numbers. Allen (1951) summarizing Hobbs and Huet's work reports that with increasing volume a decreasing gradient is re- quired to produce a given velocity. It is the velocity Figure 3 Longitudinal profile of McCoy Creek A" ‘a 74 O 720 «5.060 35000 401000 000 C 30, Get 000 \ 7 2.5 Sfigaam #64ma I.» F D00 2.0 95.000 T cofooo 5000 740 - V I if I I 8 ° 8 3 -° b- E ~c \9 S 1333 NI '(1‘1A3'1 V39 'MOQV) MDIJVAB‘IS “93mg v’z' ‘\ produced that is Ana Appears Section ' therefor tetion. the envi: of fish t No 1 Volume an velocity Acco and a Oro lineato b The dhided 1 he“histor- “at of t milewme Indiana. ”mm 11 lboy. the General d; Riv” [tya 37 produced by gradient or a gradient-volume relationship that is the important factor. Analyzing McCoy Creek first on gradient alone, it appears as though the last 16 percent (plus one-half of Section VII) of the stream.ehows proper gradient (and therefore velocity) specifically suited for trout adap- tation. The upper reaches appear suitable for trout but the environment will hold a greater variety and abundance of fish competitors than one of higher gradients. No known technique is available for reconciling volume and gradient data so as to arrive at a common velocity factor as an end result. According to Huet (1959) European's stream profile and a cross section of their valley can be used to de- lineate biological zones. The cross section of the McCoy Creek valley can be divided into four zones. The first some extended from.the headwaters, downstream approximately 29,000 feet (60 per- cent of the streamfls length). It is a relatively flat mile-wide valley, part of a prairie soil area of northern Indiana. Approximately one-half mile to the east of the stream lies a series of forested ridges 100-160 feet above the stream plain. These hills are parallel to the general direction of stream flow towards the St. Joseph River [type 3]. The (25 pert hills tc northwes in width ably [ty by very The valley c bills [15; m: the previ The waters t‘; or condu Section 1 trout Vat to treats W 38 The second zone extends approximately 12,000 feet (25 percent of the streamfls length). Closing in of the hills to the southeast and formation of hills on the northwest side reduces the valley to only one-fourth.mile in width. The current in this stretch increases notice- ably [type 2]. In the third zone, the stream is enclosed by voey steep banks 10-20 feet high [type 1A]. The last zone winds through a narrow (300 feet wide) valley completely surrounded by the now converged bordering hills [type 10]. Table 5 summarizes the survey's results obtainedfor the previously discussed topics. I The anomaly here is that the upper reaches are quieter waters than the lower stretches. This appears the reverse of conditions for most trout streams. Unfortunately, Section IX, which assays physically to be more typically trout water, is the section which is frequently exposed to treated sewage and industrial discharges. Stream.Cover Permanent cover was considered to be primarily beds of aquatic vegetation and bank shrubs. 'Uater cress (Nasturtium officinale) was the pre- dominant aquatic plant. .During the survey all cross beds were mapped. Their stream.surface area coverage was esti- mated by pacing their length and estimating their Table 5 Stream gradient and cross section types 22’ 39 Gradient stream.fall in thousands of feet Valley type cross section ‘Section V . 1.h 1.0 1.0 1.0 1.0 1.1 1.1 Type 3 Type 2 X 1.1 5.9 Type IA . & lC mar-kc: nethm dense in Sea obserw 1 Offer of at] Rated 1 cut b. as to ho approximate width. Their coverage in percent of each section was noted. Other aquatic vegetation which afforded cover in the order of their observed abundance and frequency: Vallisi- gagig’americana; Potamogeton pusillus; Anacharis canadensis: Potambgeton spa; fiymphea adorata; Podostementum spa, and 925:5. . .. Location and abundance of these vegetation beds were marked. Criterion of abundance was based on the general method in part from Lagler (1956); abundant-growing in a dense distribution on the streamis substrate, common-growing in scattered dense patches in the stream; sparse-seldom observed in a stretch of stream. Bank shrubs extending over the stream.which.would offershade were located on the survey map. Their extent of stream.ehading, in percent, for each section was esti- mated by previously outlined methods. Temporary cover included brush piles, debris, under- cut banks and logs. All such types of cover were mapped as to location. These results are summarized in Table 6. Results and Discussion.--0nly in Sections I and V were there high percentages of shrub shading (vegetation in leaf). These two sections represented the "original stream," Most of Section III was well lined with trees providing "unmeasured" shading. m:Ca£ amen mnflho H llllllll none GD smog neshm no a cmeb unshte a o owweMUH sous: Scam homfifi 3 HMII JIJV. _ anewao 22:13... e... Emfisu :5? .e pHo>flUU ll ‘ .‘u .b e 1 b a O T Percentage of cover based on total stream length and cover hl .oensasea no: .wnaoonu meal undoubown cued acumen oeo003.ha«>eem at _ .essnee oceans: as voeuue heaaaan men moo-Hana head: no wnueeaneuaa manna. H0 anlan * canebhenno “m e an H sex. one» sees a NH. nu a In cosine m m HHHb : m -- esteem ea mm HH> mach. Oflbm n me e an ceases o: on as am shaman : m # enasam sen» seen 00 > one: o m oncogene mm o >H psonunoanp Haas. m an a season m mm HHH : m ON ON HH A .eeaHH-u be: ceases» so: on ma H W -. we, 1a . w w w champ oven ouemo unsun- psoneomb omen Amman neapavoueb means. amen . .e new . eneaeeem heals uptown nonmemmumewmmmeempd anemones use: HOPOD onenesuem Th over 70 Mu pruned. with 3- vhich i devoid stream kept pr I dense Shrubs. Th lonly d Survey the Btr mid-sap 3181;th h2 The cross and shrubs in Sections I, V and VI cover over 70 percent of the water surface. Much of Section VI flowing through Buchanan is kept pruned. Some of the banks along its course are lined with 3-6 foot high wooden and concrete retaining walls which inhibit shrub growth. Section VIII is almost devoid of bank cover. Although Section IX is a natural stream section, the bank shrubs on Clarkis property are kept pruned. Below the sewage plant, large trees form a dense canopy excluding much light and light-requiring shrubs. The aquatic vegetation beds in Section IV were com- monly distributed at the time of the initialstream survey (November). However luxuriant growths covered the streamis substrate during the months from.May through mid-September. Sections III, VI and VIII had sparsely distributed beds of vegetation for 3 seasons but had common distribution throughout the stream for the period from.fiay through mid-September. In the rapid water portions of SectiOn VI, little vegetative growth was noted during any season. Temporary cover occurred sporadically throughout the stream in small amounts. Section III provided this type of cover most frequently.) ‘ The bmportance of cover to trout is well understood. Boussu (195h) found that removing brush cover along a small at fish, eV creased removal vegetati of value effect 0 effect 0 Water Ti Var lated, monly us 1956), P00 0 n) (n O h3 small stream.resulted in a decrease in the numbers of fish, even though the fish in the control section in- creased in numbers. Similar results were observed with removal of under-cut banks. In the same study, aquatic vegetation (mainly water cress) was cited to be primarily of value to fingerlings. This cover had no apparent effect on the abundance of legal-sized fish: bank cover's effect on temperature will be discussed in another section. ‘Hater Types variations in the character of the water were tabu- lated. Descriptions of these types are in terms com- monly used by fishermen and biologists (in part, Allen, 1956). ‘ Aggglgr-water depth at least 20 inches and 8-10 3 inches deeper than that of the main channel. Current sometimes turbulent, usually slight. §1£t§--Generally smooth flowing water of moderate if to slight current--less depth than pools. agar-Moderate to rapid current and fairly deep, flow usually turbulent. Uncluttered water area usually of less than average width. Riffles--Shallow water with a rapid current and usually a broken flow. Cascades-~Hater in which a steep gradient combined with a bed of stones or rocks produces a very rapid, broken flow, often with white water. These regular at Table w III, IV, I frequent 4 III). The 1 described The : deep. fair rowing ob help acco gradient The and lubst °r Substr Percent-,8E sections were comp II. III, IMS is d under the trees bed; silt is p: These water types were mapped during part of the regular survey. Table 7 summarizes the data by sections. Results and Discussion.--In the dredged Sections (II, III, IV, VIII) flats predominate; riffles are very in- frequent as are runs (except for a small area of Section III). The abundance of runs in Section V has already been described on page31» The flats in Section IX are a result of relatively deep, fairly straight stretches. The lack of stream nar- rowing obstructions and relatively uniform.width also help account for this condition in a section with a high gradient factor. The data suggests a correlation between water types and substrate. The sections with the greatest percentage of substrate in medium gravel and rocks have the greatest percentage of riffles (Sections VI, VII, IX). Those sections with the highest percentage of silt and sand, were composed primarily of the "flat" water type (Sections II, III, IV, VIII). Section v seems-t0 be an exception. This is due to the large areas of relatively still water under the shrubs which allow accumulation of silt. This is comparable to the semi-solid build-up of silt in the cross beds. In the active stream flow channel, little silt is present. Table 7 Water types (in percent), based on total stream length and area Sections II III IV VII VIII IX Average percent tOt‘I 8 #5 Water t as Sections . FIits Runs es Cascades 1% I . Not completely recorded II 100 -- -- -- III 82 15 3 -- IV 93 6 l -- V 28 72 -- -- v1 27 so 17 -- VII 10 35 55 -- VIII 9S 3 2 -- IX Sh 10 2h 12 Average in percent for total stream. 51 39 10 .5 Cor flats 81 riffles 23311 Da locatio This we through surface The tot 88 well 151il'umen 2g data 1; unit 01 greats: Proxim aVGrag. I1 Shatte: a°°t1o are“ 0. surfac. ratio ‘ trout l M6 Considering the entire stream, the data indicated flats are the predominant water type, followed by runs; riffles being the least frequent. 29.2.1.1 Data obtained for pools were mapped according to location, depth and approximate water surface area coverage. This was accomplished by measuring the various depths throughout the pool and their approximate maximum width, surface shape, and length. Their area was then computed. The total number of pools per section was also tabulated as well as the amount of their overhead cover. The in- strument of p001 formation was noted if observable. Results and Discussion.--Analysis of the summarized data in Tables 8 and 9 show a low density of pools per unit of stream length and area. Section V contains the greatest number of pools (30). This section had an ap- proximate pool-stream.surface area ratio of 1:27. The average individual pool surface area was 113 square feet. In an evaluation study of stream.1mpr0vement devices Shetter g§_gl. (l9h6) lists statistics from a 1600 foot section of "improved" streami The 29 pools had a total area of 5,720 square feet and an average individual pool surface area of 238 square feet. Its pool-stream.erea ratio was 1:5.2. Better fish yields and increased brook trout numbers were attributable to a greater survival of Table 8 Pool survey indicating their abundance, probable mode of formation, and relative abundance of shading. h? 282 0 H35 mm cm .. mm mm : HH> c on em on ma mm Ho a as co mm mm cm s o mm mm m: mm o cH. -- co 0H _ on we ow HHH essences» eoz HH . ceeessneo.ocz _ _ . v H heave adamwc cpmoo ansmnm hoboo moonnobo nHmMm encupoom meson . Ana: «doom nonasz noapsahom Hoomfi Sectior Has '2 ‘ Table 9 Morphometric pool data Approximate Average pool surface Pool/stream area Sections depth in area in ratio inches sq. ft. I Not tabulated II Not tabulated III 26 1,000 1:65 IV 22 400 1:207 V 3b, 3,400 1:24 VI 32 1,800 1:75 VII 24 130 l:h52 VIII None IX 38 1,800 1:64 young t tative Th for Sec depth a It water 1 pool f: Temgert T1 Permane Daily 1 and 5-4 Hornets] 15). 1 Mo] #9 young trout. These results were apparently from quanti- tative and qualitative improvements. The remainder of the stream.bad few pools and, except for Sections VI and IX, they were generally smaller in depth and area. ‘ These results could be expected since the "flat” water type and low stream velocity are not conducive to pool formation. Eggpgratures Temperatures were recorded daily and monthly at the permanent station and monthly at six other stations. Daily readings were recorded at approximately 7-8:00 A.M. and 5-6:00 P.M. All readings were made with a glass ther- mometer, mercury type, graduated in degrees F0 (see page 15). Figures h through 12 present the daily changes recorded for the months of October, 1961 through part of June, 1962. These temperatures g2 22! necessarily denote daily maximum or minimum temperatures. Since water temr peratures lag somewhat behind that of air, they would more nearly indicate maximum.and minimum.water temperatures. lo adjustment was made for seasonal changes in the length of day and its effect on recordings for these specific tunes. Monthly temperature data were collected during the latter one-third of the month in the afternoon, at the Figure )1 Daily water and air temperatures for October, 1961 permanent station — e _ k“ ‘ A- — — o _ \_ \__ k~~ ¥ = \\ _ V k \K‘ ‘ 1 a \\ k ‘ ‘ ' / ///// [I A/ n_ ,' ‘\ \~- .1 ‘ ‘r 1 ‘ \\\\~ ‘ 1 g\\‘ " 1. \\ ‘7‘ . :.\ xx“ . f\\\>\‘\‘ ‘ ,\‘>\ \_“\\ l -0 Eco » 5; .5350 A. __ . I___ .U #.___ .._"_w _1_. .__n __ .A-HF .<__4_w”.__ _I. ..f .fi. .:. 1....”m1_....gfl.+w .f..;w._..f I .".:..;1 +;..:.. h ...L.:....: .:Lf.w_.__:_m._:...:-_....__.:: .T; t 111.: .L::. a .mn .H.._._._w,_ .._m_.1. _ ‘11,. _I__Il.. A“...x.ln.. w; ,.,mm-.__-* ._.n.."..* .5...“ 1......1 E .... M. .1......... .._.:."_. 1...».Mr.._..lfiuv .«flxmwgwx.:...__.. DY. _ . . a _ .__.... p ...h.... _. . f1 ...__._- 1...; _.I___..;H_..~\._a,nm . _ .. .... iii”: 1......Zm.... . E .. . I. .n.. ..x ......1.. mil... 4...: _._._HI.,m . ... ... .n1.._. . _1..“ ”P .w . N .. _ . . . _+ 1. __. _- . . _ . 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'. _- .. .. _-.______. p__..._._.. -_m-.. . .-._.-.. . ._..-- ._ _ . . . ._ . .- .—._._._.... -r——.—--—.— ._y- —- W-——'—- _ -_-. ._.~y__ e v .- - n e e -0 -O~—- I F—._._ -_._ -_. _.. - - . . .. ._ . _.. - - . ._ . -.. - I- _-..-n‘ . y__._._-_—- ._r. ,_ ._- _ . s . _. , ._ . . - ._-. _.-74 L—~~———~~—-— .._ -. . .. . . .- --..__- . r-W... ._.—_---.__--- . 7 . . _ e — --.F . ‘ l I r—o——_-——-_ *9— »- _ . - . . . a- . . ._ -___1 . ._.—._....__..‘ _-.--._..__ . . . , .-.. . .. . .- -.__._.-_1 Q .—-—.~—-.—.—_...——._*_._. .- .. _ _ _ _.. , _ . ._. ,4 . .—_ r _. .- _ -----._ _... . . ._.. -».__.__J . 1 .1._._._._.._.._._...___._._.,.__..._._..__.- ._...- - . - .. .- . ._.. - I _-.- . . . . . . “_..—._.. 1 . ._.—._..— -_._.-._ .- _ ._ . ._ . .. - ._- - .__._....._. _-.._ -. . ._ ._.. . ._....— ..-._-__ ._ ._-._ . . ._- _ .. . - . _-. -_-__...__. -L-—-——*————o—-p—»~—>— ——.—- -—-...-. _- -. . ._-... . . . ._- - _.‘._..___. fl . - . 1 -_....____4 .—__-.-_._.--.._...._._..._.... - <——b-— r__q .___l__.__._.__.._.- _ _.. . . . --._._,__1 1 1 --—— _, -- _. .- .. . . _.. ._ ”._.... 1 .H—h.~-.—_.__..— ..—.+—.——- n......_.... . _ . .- ._- _-.. .— . - - _..—_.....4 ‘ 1 1 ‘_ 1 1 1,—*_..—.__._..._— ._.—_.- .._-._—.—._____.—--_o— . e —/ _.. y . , _.._ _._ ,_ __ , _ . . , ,_ _._..__ V ———f A _.- - -. ,_-,. . ._ O ._ a- . . . o ., ~l_.—..__J . ‘ ' j p..-,__. _. v -- - .-- -_ - .- . . - .-.—.---‘-4 1 —— _... - . .- o . .. .- . . - . .__.~._..-1_‘ ”—4—... . .— 1 -.__ .n ._. . . .. . . . - 7 —o-—.—-._-4 ._-.-._......-- . ._., . .. .. .-._-n_..-1 ‘ I _.._~_-.— ._-. .. ._. ._—. . _..—bba j __ .___. ,, _ _..._.._.._ ....~—_-- - - .- _ . . . ._.__.- _ .-- ._._.- .. _ ._- — p_._-. --._ . -. ._. . -.-.-_.__1 I. - . . — ..... -_ _._1 ._ ‘ . - . . , . . .. .--.__ _. .1,_ __ _.. _ - - - o - .. _. -.._. .— ._." - . ._--.-.-. 1-.._._1 _--- . _..._.. ._-..1 RE LN DEGREEEH 1 (LG "'I O / 25 20 W01 5 NOVEMBER Figure 6 Daily water and air temperatures for December, 1961 permanent station 6 - I I V v D 0 1 -._..4... -..-..--.._ -._.--- “.._1--.--...--_._ ‘ >1— .__—.._- i I l . --. . .-_-...H. , . 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'2 .5, Tan '1 '- ‘uvom 'H We. nuns u m m “name 25 2O \5 TE. \ 5 DECEMBER 1% Figure 7 Daily water and air temperatures for January, 1962 permanent station 3:]- ,.\ -\ - \“. \ - \\ ‘- \“ \‘ \§“ 1 1 i 1 1 i ! l 4/1 1 M r \w. 1. a~‘_.-_,_. _.._. -.—_———-——?__. -7 _'—-’-'—1-‘7_ . . I. -_,r_,__m A! ‘ _4 V i i i I 1 1' i 1 L v \* [_-- ._.—._..— .. 1 1 1 .E . . . . --_.--_--» 3 , ___1 1--.-._.__- .-- , .- _.-- ._-1- .- .1 :- - ”--_.—_.-- - -1...— (C a _ _ Bahhg’u ”-_.- .- _:_. -__».--1_._4 A 7" “_DPU’TQHIEifQLL ‘7 _' 17::Lfrfl _ 1 , _ ' :i’a - . 111--..-. a. .._.-_;_;_;-- _fl - - _' LU.” F. “_.-_- - - -_- ”ML- 4 l g ‘3— ' 1._._._.- 1 - .-_....W._--._.__._-_ ,.. ,---_~ . +— I I l v I I :1 l V 1. TEE I I 1 I DRTE -_. .._ ._..-._.- J____-- o _,._..... _--..~._ _.._.— - .~.....__.._.- .._1 ,_.,, 1 1 ' 3 _.-._—_..._.— . -Lifie l _..- .._.—.._..___- 1 _.,~_._. ._.—V.— ._.. -p ' I }_.___..-.__.__._..._._..\ -__-.. ._ __-._._._\ . _ .. . . .---.-_._. -._.._ -._..__ 1......1__ 1 ! l0 - ._. ._.—4.- ,-,,__.‘_,V_ -_. 5 JHNUR 91‘? N62 --. _.-_.—_..._“--- “._- '-—-_—~— -—‘--r—-?—‘—.‘- 4- ——f-r— '-’ '—*—" -" . . , ...____.- _..—.__. L._,_..__ .._.4 -._.—._._-._4._;_..7_._ _.- ._..-._._-._._.._. ‘\ .._—.._... . . r ' '7‘..- L-.._.__._-£—~ .- -_. . 7,1,.V ._ - _t.. . _ --_ l ' ‘ - , L—‘o—q—c—o--«— -— L.”4 _.._.—_..._” -_‘._.._._.--_«__.'_—.._.._ ._ ~4_»—,—._. .— _— , 77.- _. . -._—.._“.._-,_. 1 C ) 1 1 1 1 1 ._.-_~_..q_..l_ '."-’V’ 7‘.._._‘.._ .....,.7 .7.. ,_ . , . . fi:_._~_,_b____._ a.-__-__.-1--_-_. -._-._Pr_ 1.:. N g; Lg g; 15; g u: $3 115 6 1f <3 «’3 mfg ' TEmPERQTURE 1N DeeaEEs (F°) unrest 1 'a '- 'nvuu '11: In.» mun...) my noun-q Figure 8 Daily water and air temperatures for February, 1962 permanent station 54 .mN No.6. 559ng (Q 59 M FEET (“Q T105 IN _..._ lo—_- — . -L. ._-... _..—_.._. _... _..._—_..,-.—.._. - v .._-_.....— 1_._.__-_. -_.—_.._.---._. __.- v “.._.. . 1 —'-—- - _.-. .._—“_---.. -—._ - . -..—--.;._._._. _T""——’ . -._.—.__... -_.—.__- _ L..._.__.-._._. -- _.._. '__.—_.--.._._.——-__...__ ._-—.._ _- _.._-—._..—7.__.—— 9* . _.. n. .. _,. _pn... W.“ H. .-....-1w. - . fl .. __ ._ _ _ _... n _. H h_ .a w. .. ..1. .. _ U ._f ._ 4..N11‘.n.. .._ . MM .. ._.. .__." . _ 1. .w “......4.._ .._. ”f .. __ .._.“. .1. . .......M.. M” _.. _ . . I .__ . . . 1.. .,.,.1...1.... _ . . 1.1111 ._ .._. _... .. .. ..41 MW _ .. M“... _ .u w...» _.. i. . _.._. . .-. _ _.....q» _ _.. . ”._. . -mfl.._.m . . . .1 _...“ . .._.”..niu ._ .._.... . ..1..rt._..,.r .. wdecfimaiwfivmphn» ._ ._ 1.1-. _r. 7:... r ..,......L..,. .._ :-_1.1.1.,H...... _ ,, .., . .._........ . -mwmgi. _. 1 .__.......1.. , .p... ...o« m Lfih .” g‘m‘om . t -__..___..._._-_— ._-_ ._. _..._-__. _..—_.._.. .- .- --_-— _. .- 1 d1 . N l <5 m' TENERRT 1186 IN DEGREEW) .<. _. . . M _-.---_-..— - -5 o... ._.--. - . .. {integer 1:1? p 254:3 2.53:. 38‘... I. e. Figure 9 Daily water and air temperatures for March, 1962 permanent station tmv mu" 'm' u.) \"unlufl 'A '- ’rmmu ‘- Ouv—P‘” - .- 1- ._..--- 1 J 1 I I 1 M _-T_. - _.-_-_.-- .._—.._.‘—,—.—. .- _..-.__-p.—o—-—.—..— 11. ,-,,.. ---.._._;.__...1 _... .- 7 O ._ ,_ .- ..-_..-_...... “1...... -. - . . 9—..- .——- —-‘_-L-vl~ " “--_-—H--- — , ._.-..__ .-‘ -- ._ .__._._. . . .. 1 -._ .__-_- .. ._.... --1._.-._ - . 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I 1 .— Y , V w:¢__‘——.—-———--—-O—o——p— -—a———-—-.—.—a._. . _..._..-.1-_.._..__4_1._..._--..- _.._.-_....- W... 1*M_......w . 1 70- d mgr“ <3 V- T O m'dld mmNN 9' TEMPERATURE IN Damask") 91.191119:- fir“:- 1N FEET DRTE Figure 10 Daily water and air temperatures for April, 1962 permanent station \1‘11 I fix /\ 1 11111111111111 111’11/1 56 PEG N030, A $ch w; :1. -IKWIZS .__Iwb._I:I.:::I: l_..__ _.._..,..-_.Q __:. . . . l. . .: de:_::w:: ”$117 _: .1“ Z R 3‘51? __ .2. _. .. . I. _ _ j. . m _i am .i f, ,lrIi T 7 IL. 1 "in T . _ ._ m _ _ , . _. h . I/Llllr Illl). . , _ M M _ “I _ _ l m“ r Wlw ”_..P.ML H. Illa-ll. .__b. _v,_... AER-’1 ._. H4. — .u ..H l. .H.<.~_”“A_HM ._ _Mh .WN , .._ ..,. . m‘.n. ” .__ m .“~ _m . ”_ >._ ., l_ _“_ ._r ll P.‘ ._ - ._s H. r. Kill... .7»... -_.”; 7.? ..f... T. .._;ZTQ 4. if; 1 . w»... “in. .. .HM. ._ _w. .... H__»,.H._un Mm -_H..._. __ «.l ..._.__ L_ 772:. ._.,fl..f_.. m.” “.._ iii”: L H ._ .._ h. l .._” ..l .l. __ .__. _. "_m l. n. ”._.“ __ ._u _ ..,.. _ 1” 4:. .ii...i m; fixiwi 9:. M :81: _ 1:3 l.w¢2.¢¢ma£m_ ¢wfi¢3 Ill :” ::_;: “H ,--_: i __ ”#1:; : ;; f 1;, L .. ..C ._r..: .v.: ...._ _h 1.. Z. _. Mpg... 4“,. 7 ff f“ 7.; g: : _Z; _ fl”; _ .__. fipfifiafiézl: IFI. _cm ff. 5; L..W.I._F;..CL_ j_ “w .h__::_._; l “._C: 717:; If. ,LTQ; T l __:;._: _ iii. if. .m.n_ u"___." l, . “ .3:: ..u_. .m. f.._.:_..fy;:m:_:w . fii._::._:. mm :.:.:_ :f. :::f L h. " ::. .I...H_.. kw“ “h V...Pb _ NM .._ ._“.H ::.:::_._ ::::: m _ Q MHNW. d.__ .M.“” MW."_‘_ . ”T H _“ .._ ”__ m. U. _L~_ “I. _U M Z: __ we ilgflifl 7.31:: S :__.:::.u :_.yn._h “8 EB Teowemmae IN DeaaEesm $.3 9 . _ , 2 H, l _ .. .— . a . _ k ‘ ¥ . ; m . . . ., n . l m . _ . h . . » tmh _. . W . . . . _ . u w _ l . _ . . L m u . . h . . . P , * . w + .. .. fl . n . _ “ flow » LI- _ _ rr . _ w r (3.]. up ., ....8.i.¢-I. I 1... .1. :3: D. l 1. Figure 11 Daily water and air temperatures for May, 1962 permanent station MED 8g 28 Y I v--- v I mm _ _ON, m. o. . m M _f; ifif. ____,.: ; ":me "if...“ _:.:+:I-:-_._..#4_: fl..uw_-:g . _ _ n . f _.. I. .. . . ._ . .__._ l .Z _.w- .- ._ I. --_ .._. .._. ...- . l f; H ::: “:Et: 5:5: ;:_;.-_ it} 3:5 _;.m.;:..N.m ._.: .._f:_f.g:_fi331_.f:;;_t3.-f..::.:-; mr .. Wyn...” ..mm .__. . ..l,._ ._H._l... __.n.u . ._m__.m-_ n“ Mm_ ..._. .4 _.M.._Ww ._q hm” .W_ . _h . ”I flu“. In}. M. ..W will: fm:__ _.....MH-H . . »_.M, _. fl. .. ..._. . ”Ll. _.N. _.Mlkm .>W_I>Ira-h “ W .F . . ._.r m ._._. W“ “_hw . .hv ._I21 4%.. 3:: SE 1.... :1: i. -_ IJTM w ,_~ ”_fwf 1.6 ¢ .._._ _..._ 4....._-__. _.— .. a—I. I . -.——.‘——...-..... ;._..._ .._- pm,-k.._. . ._ ‘._,__‘.. . r ‘ C—»o——~o—~‘-o‘—.—. rp-w—fq .._- .— l I l - _.. - 5.... Y"- .. _At— ._ ._ .__..__o_._.. ._....— . O 57 -«. -._.—.._ A . ---._. .-. --.._ V. {E 53 r . if. tn .9 ..-_-. “._...— - . H. . . H ~ -‘ l .l. .._...._,__‘ .-.. I ‘ . . u I \ f a o h TEMPERATURE 1N DEBREESW) .M . m _-.? .- 3 _ _ _T .P i w I ; LN“ w l 26w _"_ I L—o-w. ..— . A v I .___'_'y..‘ H‘ .— .__ .._. ._.- _.. 6... .8 f: .— --_.-- -__._ L_—..——--__ ._- .._..- .-_.—._.. I——— - _-.. -_.___ ._ - .- _..—._....~ _.. F... .__” I._...___.-r.. -- ._. ._ ._...- ,_*___._ ._ ,_.—.— .. __ _--_-—_..._... .r_.._-__”.. y—_.___ "I. ’F a... 3!. IDs-II. '.l .0... :3’). I. a. Figure 12 Daily water and air temperatures for June, 1962 permanent station 58 v ”rmv~-v*‘v I o I I l I l I I ,' ' l . I I , . l . . L. . . o . . ' . . , . ' . , I . 1 _ .-_-— # - -._.—_—-.— .._.—P—y—.- -—-—-v——.— '— -.—.—_§—..—J . .__ _.. ._. . _.- - - -- _-._.. --_.—..-,-..._.._ l l _7 _..._.__._._ .__ . P#‘.__P_.__ .._... .... . .. .. Y A .-_ .__- _ - .._ . -_4_.-.”_ -__...U-.__ .——>—.—-—-< -._ .._.... ._.___..- .-_.. -.. _. -..—IF" __ _..... --. ~——‘—O——- ._.—.._. .._.—.._.._ I . .7 . - . - ..--.. h ..a... . I ->—..—-—-————.."—..— _- -o .7 -7 a 1..— . ». -._. - 7"- _, _i ”cap—.-_..q» --. —--‘ I-- - . l .. __ _..—.-I—“vre- I I l l l I I l l I I l l ; I I l l ‘ | l - 7. a . .-;__'Y_-_+_._.-._*.l ‘ I l ' I .4 .. . ,-.. --.—.._ ....m .._.—y... i I I I I I I I I I I I I i I I I l ' I 1:; TO I l T l l l l l l I l l I R _.—.._._._ ._. _..- - .— ,_. _ .-_.. _-b-~“’.._ ._.- _... A-+__.. V_m V*-R¢-_,_. 57.. .. - .v-. I I l U! I as I I I__.,.___ __ - _..._ -___ _._._-____.._._-- .__--- _-.- .__- .._-. .._--.-... _ .__-..- , -_-“...HH. l———__-I_— _.._____F_ ._.—”-__..-”d .... . . _- . . . .. ._-- - . ._ .- ._..._...._._,....__ I I I I l I l l I I I I Y" I I W mPE: 2 ; II I I I I II: V...” ..._'_ ... _. .. -._..._- _”,,_..... _ -- ._.... .__- _--. r- o-.— ._¥r_.7,.. -_ .__. _.‘_ _.._..I._ .__-._.-—. ._ ”h_. _..4 __~ .__—__ ._.. .._ .._...- .-._, ..V--7 ., .__v._-_ ..--_-_.-_.---- ._ ._.-_-.- ._.._._ —————--. .._- -_._ -_.—.__- ._ 7-.---P _...__- _..._”.-. _. .__ , . I rI l _i I I I I I I I I I I I I . 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I ‘ ~ _-.. ..--.__..-..._-_.-.__.._ - _-._.”..- _-._.—.__-_.._.._..-. - .>-;-...~.- 7H,";--j_..---__. __ _ .._—4. ._. a..__‘._,. ..-.7 . .-_ - . ._. .7 . ._ _.. __ - _.. _ __ ,_ - A A. . ._.— —-L-- I ‘ I ._. .-_.—---- ._.— ._.a_-._-.._ .--..4- -_ _.._-—._.. ._ ._ "-’M'v‘ -._.___ .._.__,_._._.- F _-._. .__.....,.. .- . .__..._ ...d .. . -_.-_._.-.._ . . I l . . I . I . I A -—*~—p—o—*—w—.——.—-_~ .__— n._.___.._._4_....._._. .-...._.._.v_.-_.. ._-A._-. .__ -_-_k_-, . ...._._,_.. _.._. -_...__ ._ L_... -_.- ..._ . -. .- .__... -~:— -—<——-l— - l l l I l l l I l l l l I l I l l l l ; . I L . I l l l l l A ‘. — .._.—...- _.._.. _-.—._.“...— V .-_—_.._...w... .__...I_.h__.._.;__.- -._.—“#7 _-- I— .___.~, . . . .._. -.-I .-.--N.._.__a l . I . . I A A g I ...—._..— .__..__._..__.._-._ .- -__..-._....l ._. _.. -_4‘ ._ -.. .._ . , . . I-.- _ - _.I._._ -_ . l ' I Y7 — .-.... --—-»o— ..--.__._ -fi.. ._.—.._.... .— ..._--_-._ ...... .- p.__._—— ,— .._...-.. -..—.__..- _.-—.._.---“ ~—-—— 0 -—-— ~—- - I ~ ,. ~—‘ —-—- D ...H. . .. . ‘ . ‘ -~ ---—~,——~ .""""“ vim . . . . . 4 . . . . J O- '5- O- 5- O- .5- LL .p-Jv IAIA'U‘I-on I‘Q' ‘rium'o to mveemues LN DEeaeeem N FEE '1 '- 'IJI-In 'II 59 stations designated in Fig. 2. If possible, days were selected on which the air temperature was above or below normal. It was postulated that the moderating effect of ground water could then be detected more readily. Figure 13 illustrates graphically the data collected monthly for the 7 stations. Results and Discussion.--There seems to be only gen- eral agreement in the literature regarding habitable temp perature for brook and brown trout (§glmg trutta). Embody (1921) stated that brook trout prefer waters with.maximum temperature of no greater than 70°F., optimum for growth being equal to or less than 70°F. They will tolerate 75°F. water under certain 02 and 002 conditions. At 83° heavy mortality occurs and produces a deleterious delayed action effect on the remaining population. Brasch‘g§_g;. (1958) summarize the literature, stating that exposure to temperature above 780 for more than a few hours will kill brook trout. Fry (l9h7), states that increased temp perature above766o results in decreasing brook trout activity; this activity declines until lethal temperatures are reached. He found that brook trout could withstand 77° temperatures. ' It is recognized that brown trout will tolerate and thrive in warmer water than brook trout. Embody (1921) refers to optimum temperature as equal to or less than 75°, but growing in waters which reach 80°F. Needham.(l938) Figure 13 Monthly air and water temperatures from seven stations 0 0 F0 -\) qo’:::0£;-h LEQEND' " ‘ ‘ ’ I" “““ ”ff—”W‘- _-.-“ ”WW?“ :ifiegigiaRwCa-TER:--"-~‘-“'1' :: O LUHTER TE mp E a F-‘ITURE .D ,_.7_._- - -_.__._..__-_.__+ - ...__.__-.__.._____..._J ._.__. -_.—_--_. --_.—._‘_-_.—‘—_—.~—_..1 _._- . .._- .._--.. - ,4- _._ _ y- _. .._.—.__q _. ... _.- _ __ —__...___.._._“——.o—_. . --_.--- . _.-—.__-.. ._.—_... —w_.——_._——-—¢ nu!“ 61 also found the 75-800 range suitable for the growth of brown trout. He gives 81°F. as the maximum safely tol- erated temperature. An examination of the daily air and water temperature data for the permanent station, shows a generalized pat- tern on warm days. 'When the recorded air temperatures were between 75-810, the water tenperature did not rise above 69°. When the daily air temperatures exceeded 820 the water temperature rose over 700 but on the hottest days, the water temperature did not exceed 75°. The morning water temperatures following the previous day's high air temperature always declined to 55-650. The warm months then, showed an approximate 120 difference between afternoon air and water tenperature; the water temperature not surpassing 75°. Two spot checks in late August, during high air temperatures (90-950P.) showed a water temperature no higher than the same reached during 900 weather in May. The nininnn.water temperature during this period also corresponded with those during May. The significance of these high, late summer air temp peratures appears to be that the water does not store up much heat energy as the summer season progresses. This suggests that the influx of cool ground water is sufficient to offset stored heat energy in the stream. 8 Eight days in May, recorded with afternoon temperatures reaching 85-9h°. 62 While the air temperatures in January varied con- siderably for different periods of the month (a 510 variation), the water temperature only fluctuated 13°. The lowest water temperature reading was recorded at 33° on a day when the air was ~120. Monthly winter water temperatures for different stations, showed a gradual decrease in temperature as the stream flowed towards its mouth. Streamredge ice increased in area downstrean.from the permanent station, reducing the main channel to a 2-3 foot width. These lowered down- stream water temperatures and increasing ice coverage might be correlated with minimal ground water activity. More seepage apparently is present upstream from the permanent station, thus preventing stream-edge ice from forming and maintaining a higher winter water temperature. Except for the headwater area, the water temperatures during the warmer months show slightly cooler temperature downstream than those of upstream stations. No definite explanation can be given for this apparent anomaly of summer-winter temperature reversal. However, one factor may be that diffuse ground water supplies are only active as they diffuse from shallow surface seepage. When the ground freezes, this seepage stops until a general spring thaw takes place. The lower stream sections may be retaining their coolness, in part due to better shade conditions. Table 10 Table 10 A comparison between water temperature gradients and water shading gradients have been computed using Section I as the base. A + number indicates the number of degrees higher the water temperature is compared to that found in Section I. Temperatures are from stations located near the end of the Section listed. 63 .H> coupoom no one Beau pooh mmw haeaeaaxonmms cenaevno moauasnemaoa ** .moeAp hp conga nofiaeen mane * m+ m+ 9+ m+ o m- oa m aaHHH> ~+ m+ m+ m+ o m- a: pH HH> ~+ m+ oa+ m+ o m- on o: H> nu nu u: an I: nu mo m >. m+ oa+ HH+ m+ m+ H+ mm o .>H 5+ oa+ oa+ m+ m+ o m: m aHHH s+ mH+ HH+ m+ o+ H+ o: om HH 0 o o o o 0 mm OH H oeuaea uueaueea pmsws¢ enabl. bamF anam¢ none: .hsuanmmh. sputum enshmm apneaoenw chapeuegae» nope: manages ”neat oceans noapoem «one eeahnnh aemsfirmo ewenneenem 61!- compares monthly water temperatures with shade development. The headwater branch is arbitrarily given a zero value against which other water temperatures are compared. The dif- ference between the base (0) and the recorded temperatures is the temperature gradient. These gradient values are the highest for Sections II and III, decreasing with increasing length of stream. Comparing these gradients with the per- cent of water shaded and/or covered, two things are sug- gested: 1) that the stream.ehading helps suppress maximum daytime water temperature; 2) the greater the degree of water shading, the slower is the seasonal rise in water temperature. By.August, the stream's water temperatures reacted quite uniformly to high air temperature except for Section I. This data suggests the importance of stream shading to water temperature. Examination of the temperature data for McCoy Creek indicates tolerable temperatures exist for both brook and brown trout. During the hotter days, brook trout may congregate in the vicinity of spring water sources (Brasch g£_gl., 1958), deeper pools and possibly in cress beds. Spot checks showed these beds to be about 5-60 cooler than the main strewn. Brown trout should find the streamfis temperature suitable for a high physiological rate of activity. Temperatures are higher than ideal for brook trout. Water Analysis A routine water analysis for various chemical cro— nerties was conducted during Fecember, 1961. Samcles were collected at eight stations and tested for dissolved oxygen, free carbon dioxide, nhenolnhthalein, and methyl orange alkalinity. The oxygen content was determined by use of Alsterberg's modification of the Winkler method, described in the tenth edition of "Standard hethods for the Examination of Water and Sewage" (1955, no. 250-251). The free carbon dioxide content was analyzed by the methods described by Lagler (1956). A summarization of the data is presented in Table 11. Results and Uiscussion.--The stream was found to be ‘I nearly saturated with dissolved oxygen at tne stations sampled. Carbon dioxide content varied between 5 and 9 com. with the execution of the water taken from the last samoling station; here it was 1h com. The then nartially treated sewage, discharged near the last samnling station, may have had some effect on the oxygen and carbon dioxide content further downstream where oxidation of the residue would be taking place. At this site a some- what oily, grey discharge was beind circulated into the stream from Clark's factory. This discharge aooarently did not affect the oxygen suonly but may have been in cart resoonsible for a higher carbon dioxide content. Table 11 Summarized data for chemical analysis of the water 66 .Afiaesv nope: amazon one hnopoeu Soak upsesahuo on 050 seen cenaea snapshoasep nope: .henw aoHoo acne: as .noHoe nope: :« emceno chansons onunaeeape can“ oemasnemac wagon usesanhe 09h» snow m: .ose a tone mm mm 0.0 0.:H +ooH m.ma m: mm em\ma seem mm mm 0.0 o.o ooH ~.:H 0: mm e~\ma aw am an 0.0 o.m ooH m.ma 0: mm em\ma smm m: m: 0.0 o.o OOH m.ma mm mm em\ma mm o: o: o.o o.» co m.mH mm mm mm\mH ma mm mm 0.0 o.m on ma on em mm\mH e H: a: 0.0 -- no NH H: mm Nm\ma m H: a: 0.0 0.» mm 0H a: an .mm\ma .m 3mm. Ema Ham Ham m _Emm meomwec moeawoo Hausa Amooaov aces soapA one» one» area .on haananexae LII nunsuem neaemaep neaouEe» even soapapm amasaaasHa Hamoa 67 The total average alkalinity content (he ppm.) ap- proached the minimal average values (h5 ppm.-200 ppm.) for waters in mid-continental United States (Ellis gt_gl., l9h8). This relatively low alkalinity value may have some biological significance. Taft and Shapovalov's (1935) general findings indicated that a stream's richness in bottom fish foods are generally correlated with high alka- linity reading. Armitage (1958) concluded that for the Yellowstone River, CaCO3 alkalinity appeared to be the main factor in determining total quantity of stream organisms. This topic of productivity‘will be discussed in the next section. Biological Conditions Objectives A survey of the stream for biological information included: analysis of the streamis bottom fauna produc- tivity in relation to the substrate and aquatic vegetation; obtaining data regarding the fish population, past and present conditions; surveying the stream's potential spawning ground areas and their relation to ground water. No sampling of the stream's plankton was undertaken. Bottom Fauna The objective of this phase of the project was to obtain an overall picture of the stream's production of t‘ 68 benthic macroorganisms, especially the insects. Further comparative analysis of these data would aid in deter- mining the ecological condition of the stream. Although total agreement is lacking, late fall and early winter appears to be one of the periods of greatest immature aquatic insect abundance (Maciolek and Needhmm, 1951; Ellis and Gowing, 1957; Armitage, 1958; Needham, 1938). The samples for this study were collected during mid-November, 1961. Needham.and Usinger (1956) demonstrated that for one riffle, 2 or 3 samples were sufficient to be reasonably certain of obtaining representatives of the principal groups of bottom organisms present. From.this, it ap- peared that the samples from the 27 randomly located stations would contain representatives of the prhmary groups present in the stream.eubstrata. Using a Surber sampler and the method described by Welsh (19MB), h9 bottom samples were collected. Unless the substrate appeared very homogeneous, two samples were taken at each station, as suggested by Lagler (1956); one each from.the center and west (or north) side of the streams The material collected was washed in the stream while still in the net of the sampler. It was then placed in a Jar, labelled, and later refrigerated for future use . 69 When the material was to be sorted, it was emptied into a large, bright aluminumsbottomed pan, then flooded with a sugar solution. This modified floatation method is described by Anderson (1959). All insect fauna and crustaceans were removed and placed in a 5 percent formalin solution containing borax. Most mollusk shells were empty, therefore, none were enumerated. Two separate samples of each of the four common types of aquatic vegetation were collected. One substrate sample also contained mostly Odostemumceratophyllum.9 Approxi- mately a one-half square foot area of vegetation was up- rooted and bottled for each sample. Later, the plants were washed thoroughly over cheesecloth.which served as a net to catch falling organisms. Identification of all fauna was accomplished with the aid of Pennak's "Fresh‘Water Invertebrates of the United States" (1959). "The insect identifications were checked by Dr. G. Guyer of Michigan State University. A synoptic list of all organisms collected and re- corded is presented in Table 12. Results and Discussion.--The data collected from sample analysis showed Tendipedidae (Diptera) to be the most numerous insect type collected. Cylloepus (Coleoptera) was the second most numerous insect while gydropsyche 9 The substrate sample was taken on a flat rock. This plant comprised most of the material sampled. Table 12 Synoptic list of bottom fauna collected 7O N oooHnoHBm onoHnuooz OHEopnOHHum H masopsowHHO H onchmoooohm m N H H N osHHmoanHH Hm : a: m n O m onossoosesm «EomoueoHo oHHnaooohnm m m H oHHeHm oooHuoaopooHom H H H oooHHHpoo H onoHnoopoSom H ousqnoH nonH oaoooaoum oHnomoKom cacaonam adamaeon «HpoHnmopooHoaom H ospoHoE¢ quoem H oaoumonesonmm N N H H : usaoafioo oHHoHohm usHHom< oonHosaHm oecHoHansa oooHHSoHaossq H H H H H oeoHcon m s H a H H osOHpasonHoo< m «coanoomHHO mm OH mH NH OH mH lawn mH NH HH OH O O m m m m m m H sHossm LH OH 0 m ..Hli m m m m H compepm In HHH 71 cocHHOHEm oHnoHapooz oHaopooHHum odaopmowHHo H H onothOGOhm H H osHHnnonfiHH HN N onohomonphm «Souomoon H «HHnoooaaom uHHon H oocHuoEopmoHom o oooHKHaoo ohoHnoopoaom oasnnoH aoaH N cacaououm oHnowoKom ohofionmm osasnnon oHnonmouooHoaom edueHoB< oHpoem anoamonoaonam m o N N N m m H condense H H H «HHoHasm H N e«_HHoeHH oomHosaHm oooHoHHHnaa N N H oooHHsoHnnssq N N H N oouHoeHn m oooHpoaouoHood spoonoomHHo MIA r4FHDCU bud «nu ._. N :Nm om +Wm mm mm mm Hm Om om mm mm mm mm mwmnumwunmmlaww oHossm H H H aH MH mH - noHeasm 72 .OOO.m: a oonHHOHEm H «HooHscooz H uHsosmOHHem oHSoumowHHo : m emomemocohm H s : ssHHeoossHH : om m N onohnmosohm H N oaoooonHo H sHHnooosanm : mHHaHm oocHaoaoamOHom O osOHchoo H H ohoHnoouoaom N am asscemH H nomH N N wH : osocouopm m N oHcomoHom m oneEonam H monopoon H H : sHooHnoopooHecao H N secoHos< H H sHpoem H ohopmoaoEonmm on m: ON NH H m H H H m accuseso HO NHN H «HHoHssm NH O msH H msHHom< N m N N socHessHm a esOHoHoHnsa osOHHsoHsossH m N H HH 3 H O m mm onHOst oooHpoEOmOHoo¢ ovoonoowHHo _ImmIIMMIIHmIomlo: may we mmllm: mm. m: N: H: on om mm OHmsem em eN mN mm: mm ,NN cOHmaom H 73 OOOHAOHoow osOHHssoesc< N H N Omoohano H H escapee H osOHHasoHsOopO H eoOHaoonOpohoo NH O OH ON m N . a H .N e N N osOHOoOHOcoe N m oOOHHHaeHm H ooHsaez oHcHoom onoopo< H OquEHH oHHnaonfiHHoosoem m OOOHHsoHa onopoHn cHHoosaoHoo N oOOHooHom N osHoHHom H OquoOoama osnoHHom H . N m H usQoOHHho oaopoooHou ON OH Own Oar OH mm (Ow. mH Nat HH OH O O N O O m, N H oHOasO HH Owl O, O O! O (m m N .w, qusssm 7h N ms H m: m OON OO O: HH m H a :H OH N : Om HN HH HH oooHavwnmm oaOHHasOnsoo amoebano assumes osOHHasoHssssO ooanowomoposoo oechoaHonoB oeoHHHsSHm ouHshoz chHoom snoopa< oHooEHH OHHnoonaHHoodonm oooHHsmHa enouoHQ «HHoesaoHoo oocHooHom osHoHHom oschooanB monoHHom osmoOHHho unoonoHoo Om HN Om mm in mm (NO Hm Om ON ON ON 0H mH 1OH wH ON ON ON ON ILHH OH MN NN HN OH NH onmom noHpoum 76 (Tricoptera) ranked third. Samples containing crustaceans indicated Asellus (Asellidae), Hyalella (Talitridae) and Gammarus (Gammaridae) to be the most common forms (listed in order of occurred frequency) in the collections. In Table 13, the amount of substrate types (in per- cent)10 for the whole stream.1s compared with the pro- portion of substrate types (in percent) for all samples. This comparison indicates that loose silt and sand-silt types were undersampled. Sand and fine gravel-sand mixtures were oversampled. Medium and small rocks were sampled slightly disproportionately to the stream's substrata com- position. Judging from the conclusions of Needham and Usinger's (1956) riffle study, statistically significant productivity data for McCoy Creek's substrate probably would not be obtained from.h9 samples. However, a food grade scale (Lagler, 1956) was used to arrive at some basis for com- parative substrate and vegetation productivity. Figure 1h compares the bottom type of each sample with its calculated food grade rating. Each bar, repre- senting the degree of food scale rating is divided into the total number of organisms tallied and their volume (in cc.). Data from Figure 1h is further broken down for comparison I 10 From Table 2. Table 13 Percentage comparisons of the stream's total substrate composition with samples from different substratum 77 Loose Sand- Sand-fine Medium Small siltfi silt Sand gravel gravel rocks Composition of streamls substrate-- in percent 25 32 h 18 8 5 Composition of substrate samples-~in percent 16 25 16 28 12 2 * Silt not in water cress beds. Figure 1h Substrate samples in relation to food grades. Vertical bars in the food grade scale are divided into: bottom half--volume of the sample (in cc.); top half--the number of organimms collected per sample. Vertical substrate bars represent the principle soil type of each sample. S: L b a: UUIJ._"JIJ\JJ. ‘ I '.I sup,- 2% 161713 {9 26 Z! 2 OUUO'JUOoOOoo-e'oc 0 Q 0 v a‘ \ \ . \ . , . , \ \_ “ > , ~. \ y \ \\, e \ -- \ \N \\\ V _. \H. V \‘ _ . ‘ e ‘\ \‘ \x‘ (55s send-5mm some m 0: Rue CARMEL a madman mm ROCKS 83 F090 (..ng 79 in Table 1h. This table indicates the number and percent of samples by bottom.type, in each food grade. Discussion of Productivity Pennak and Van Gerper (19h?) found that gravel yielded two and one-half times more bottom fauna than sand per unit area. Shetter g§_gl, (l9h6) noted that as the current exposed additional gravel bottom in the stream, the bottom fauna population eventually increased in quantity and quality. Morofsky‘g£_gl. (19h9) mentions similar results as does Tarzwell (1937) from.his studies. Analysis of the data suggests results concurring with the literature regarding general productivity. Sixty- seven percent of the medium gravel samples and 100 percent of the rock samples were in grade II. Silt represented 16 percent of the total substrate samples of which 12 percent of the silt type was in grade III, 88 percent in grade I. Sand-silt comprised 25 percent of the substrate samples of which 8 percent was in grade II and 92 percent in grade I. These results tend to agree with the summarized findings of other workers (Murray, 1938; Tarzwell, 1938; Tebo, Jr., 1955; Cordonc and Kelley, 1961), which indicate that sand and inorganic silt substrata were very low in bottom fauna productivity. Table 1h Number and percent of samples by bottom and vegetation types in each food grade 8O nu un nu anomaom munhaauhonow canamm msH noeazmnnhaaaeoa pnuunsn< none oaspom nn un uu anomeom mnuhaaeooa noafioo esooHaoSo aoaaamnuhaaeooa anavnsnd museumHHHa> nn mooa u Amy nu haaaneuem mmncemaneo oneness Haannennamm uaamnoand .. mom n AHV mom n AHV mangoes owsswoacoo unomuom HHuuunoHEoo asfipnspamz mam n Ao:v «as n Amy mm u may «suspense Has sou seasw eoou . some a“ paeohem and 9035:: Hence . nu Rooa n AHV nu N exooh HHeSm umm n Am. mac n Any un NH Hopasm asses: Row u ANHV Rda u ANV nn mm Aeaauvaebaaw . one oaam mooan Amy nu u- on enam mwo u many an n AHV -u mm pflaa\eqsm puma n Rev nn : .. mma n Adv ea anew H sumac room HH ooahw pooh HHH enema coom _ mome epunpnnfiu can“ conEam noose coon pceaonuao on» ma moa San Apneoaem hnv oopeapaeneuuac soapwuewep openpuASm ho unooaem one hopes: aeuoa nonEem Hence no euanpnpsn _mo e93. 81 The one silt sample in grade III contained a very high estimated11 number of Oligochaetes. It was collected from.substrate in water subject to industrial and sewage wastes and far enough from the waste discharge source to allow the material to decompose. Goodnight and Whitley (1960) state that it is thought that Oligochaetes may serve as indicators of pollution. Two percent of all samples were in food grade III, 17 percent in grade II, and 81 percent in grade I. This data suggest low substrate fauna productivity.12l ' An analysis of the vegetation sampled suggests their importance for harboring aquatic fauna. Data recorded in Table 1h show the 2 cress samples in grades III and II respectively. Both Anecharis samples were in grade II. Vallisneria and Potamogeton pusillus did not yield any organisms in November. However, visual inspection in June of these two submerged species, as well as two other species of Potamogeton not present in November, showed lgggg_numbers of clinging Simulidae larva and cases. This food adds measurably to the total summer's standing crop in the stream. The presence of a few species of stream vegetation 11 Number of organisms in one square in, counted and total organisms estimated on this basis for the rest of the sample. 12 Prom.rig. 1h; 87 percent of total stream substrate in sand mixtures or silt. 82 during the months of June through mid-September is very noticeable. However, except for cress and Anacharis, the volume of vegetation declined drastically during the months from.mid-September through.Hay. As noted, Anacharis is scarce at any season. Water cress, however, begins ex- tending their growth during April and reach maximum growth in July, which they maintain until approximately October. The beds then decline gradually until a low point is reached around the first of April. The limited data collected suggests agreement with Lagler's (1956) summary of Needhamfls (1930) work, indi- eating a seven times greater productivity for stream bottoms supporting vegetation than substrate barren of rooted plants. It must be remembered however, that com- parable productivity of vegetation and substrate in this study is difficult since the quantities sampled are not comparable. 8 It appears as though aquatic fauna samples for McCoy Creek would best be obtained in early spring prior to hatches. Bottom Fauna and Velocity ‘ Both.Rees (1959) and Curry (19Sh) found that the principal factors affecting the distribution of insects in a stream.eppeared to be current and bottom type. Rees found Diptera to predominate in slow sandy bottom.areas; Coleoptera preferred gravel, Ephemeroptera, Plecoptera and 83 Tricoptera, swift flow and gravel bottom. His data does not include possible variances in substrate preferability based on insect generic differences. Curry's work with Tendipedidae found individual species were restricted by current and bottom types. The data in Figure 5 portrays the relationship of velocity and substrate to insect numbers classified by order. The predominant insect genera comprising these orders are listed on page 81. Based on these predom- inating genera, the greatest number of Tendipedidae (Dip- tera) and Hydropsyche (Tricoptera) were found in gravel substrate and slow water. Cylloepus (Coleoptere) was common in both gravel and sand~-fine gravel mixture, ap- pearing to show a preference for moderately fast water. Seventy-eight percent of all samples (38 out of h?) were from slow or moderately slow water (this water type com- prised most of the stream--12 percent (6) from.moderately fast water, and 10 percent (5) from fast water.13 Six and three-tenths times as many samples were taken from.slow water as from.moderate1y fast water. For a better comparison, assume the insects will be taken from the moderate velocity water in the same proportion as the actual sampling. Then 38 samples from water of moderate velocity might yield approximately 6.3 times the numbers already presented or: 13 From water subject to pollution. Figure 15 Velocity and bottom type relationship to insect numbers--by genera 8‘} .53 33523 on mm o N H NM n Ema amok—805:8 m < E02“... U {dmhoowmou g mamas Home” “a .5 .3 WE waging a 5%.; 923 mm 3 S b . gun Che. 92%. w .5295 wzfi .3335 .95 PF .32 - v 0d” 8 _ . ~ 3 . . ..t u_ 3 mm flw w an , w , , - inn: nan , W N a: \\\\\\\\\- u an. a I an I m. to n H mm W w. ...A d . 3 B O D. .mmmééiou .52 $.45 unazqw totem - ..zofiajoo . c» 52.96 mPEB 2. 92523 - 333/11 9."! C 85 662 Diptera vs. 319 for slow water in gravel 13 " vs. 151 " ' " " " gravel-send 32 " vs. 36 " 8 " " sand 3h? Coleoptera vs. 21 for slow water in gravel h79 " vs. 7 “ " " " gravel-sand 6S " vs. 16 " " " " sand 227 Tricoptera vs. 55 8 " 8 " gravel 32 O " vs. 67 8 " V * gravel-sand 0 8 vs. 12 8 8 " * sand The actual numbers have no significance except to show the trend of environmental preference by the insects concerned. Trout Food Preferencgg Another method of evaluating the food value of a stream.is to consider its faunal production in terms of preferred trout foods. Although no stomach analysis data were collected, it appeared that a comparison of preferred trout foods with a sampling of invertebrate fauna from McCoy Greek might be a useful evaluation tool. Judging from stomach analysis data, Horofsky (193k) found brown trout to prefer Triooptera and Ephemeroptera. These two orders were also the most commonly distributed groups in the study area. Ellis and Gowing (1957) stated that where plentiful, crustaceans and mollusks also seem quite important in a northern Michigan stream, Brook 86 trout in a southern Michigan stream.elso showed this food preference (Morofsky, 193k) as did larger browns in an English study area (Frost, 1939), Shetter gt_gl. (l9h6) quoting Leonard's findings stated that apparently certain species of Tricoptera, Tendipedidae and Hyalella (Tali- tridae) were favored by brook trout in Hunt Creek, Michigan. A western United States rainbow (S3122 gairdnerii) and brown trout study (Maciolek and Needham, 1951) indicated the importance of Tendipedidae; being preferred even though low sometimes in availability. Although sampling data based on a food scale rating indicate low food production values for McCoy Creek, two out of three most common insect types sampled seem to be good trout foods. The same type fauna (Asellus and Hyalella) harbored by some of the vegetation in the study stream.eppeared to be readily eaten by trout in other Michigan stream.studies. Since water cress,anacharis and other aquatic vegetation seemed to provide habitat for a greater population than did most other substrates, any change in the volume of vegetation present in the strewn may play an important part in effecting the stream's faunal population. Fishing Verbal information from the previouslyrmentioned cooperators aided in obtaining a generalized relationship 87 between trout fishing success and changes in stream.ehar- acteristics. _Originally, it was reported that the stream offered good trout fishing throughout most of its entire length (6, 8, 11). However the area flowing downstream through and beyond Clark's property has been and still is, subject to industrial wastes and to a lesser extent sewage dis- charge. This waste has ruined all aesthetic values of fishing in these areas and may have been responsible for a lower catch (8). Stream conditions were different prior to 1935 than these existing now, and have been previously described. Prior to 1900, bluegills (Lepomis app) and bass (Micropterus app) were caught in the stream below the mill pond (Section III) dam (9). Reports from the years after 1925 indicated that the most abundant trout species caught were rainbow and brown trout (2, 5, 11). Brook trout were taken from certain areas throughout the stream.but more commonly upstream from M-6O (8). Limits of 15 rainbow trout per day were caught frequently (3, 8, 11). Summer fishing, using grasshoppers as bait, was very good (5, 11). Fish size varied in the catch from less than 7 inches to an occasional 16-18 inch brown or rainbow trout (3, 8, 11). It was the consensus of the cooperators that undersized trout of any species had not been caught for 15-20 years 88 (5,8). However, very little fishing has been done by any of these people in recent years. All informants (3, S, 8, 11) emphasized the fact that fishing was very good in earlier years. As physical changes in the stream occurred, smaller catches became the usual. Fishing interest waned. Prior to l95h the sections north 6f M-60 still provided fair catches of brook trout (8). As this stretch became silted in from.ditching operations, the fish yield--and fishing interest, declined. The area from the railroad grade downstream to the city limits of Buchanan may still hold a fair population of fish.with some large trout (5, 8). Much of this area is almost impossible to fish in any fashion due to shrub tangles covering the stream. 0f the "non-game" fish, suckers were the most abun- dant, especially during the spring. Presumably they entered through the Pike Lake drain, spawning in the creek. Horny head chubs and grass pickerel were frequently caught (5, a, 11). Although the major fishing effort is apparently di- rected towards catching trout, other species are sought. As in former years, suckers can be found in the creek during the spring. At this time they seem.to be present in catcheble numbers and sought by younger fishermen. Bass can be taken in a small dredged-out connecting pool It. 89 nonopeo poems .psoa ea pounce arena .FH .maa Macao have: .Apnwaav ocean use Anneav tonnaea one each» peemasa o3» one .msoa as nausea paces .ea gnu. ‘_?. . IA ‘ \ i p - X . t. . 1 . 4 \. 'e n n . . a a s _ .§ . n u u.. .l ,. . n _ ‘ , . e no i: .n . \r . .e u x o e o z o c. u 8 5 n O H '1’ ‘ ' I‘I u, I . ’n r l - 8. n I _ n ' Q .. u. n/ Is , .. 1 Yin . \l .1 . .|r n. b .7, .u . II On no P O u n.’ O u (e'.. .Uununln .n I'. .7». .a ‘0 u NH ovwdm r: ... Brew. ,. Mia... .... .mam mu ae.,- t \ I 4. . N c . 3&1. ’ If... 1 $-51nln \ I lull ‘ I! 90 Plate X Fig. 18. Trout caught in 19Sh from McCoy Greek . ll! |[l‘l‘ 91 one-fourth mile upstream from the Bakertown-Buchanan Road bridge. These bass are only in evidence during a certain time in the spring. This may correspond to their spawning period. Other warm.water fish.may be present in the two ponds in Buchanan McCoy Creek has been stocked1h with trout by the Con- servation Department as early as 193k. Since l9h8, the department has, as policy, managed McCoy Creek as a trout stream, Table 15 lists all trout stocked since l9h8. A total of 1,000 legal size (7 inch) rainbow and lh,h00 legal- sized brook trout have been planted--no rainbow stocked since 1955. In 1959, 20,000 brook trout fry were released as well as the larger fish. These additional small fish represented utilization of a hatchery over-production situation. The 1961-62 program called for two plants of 300 each; one just prior to the opening of the season and the second during May. Each plant was divided equally between the Buffalo Road bridge site and the Bakerstown- Buchanan Road bridge site. Fish size in the first plant was between 8 and 9 inches; those of the second plant were slightly smaller. All were over minimum.1egal size. None of the fish were marked for future identification. In All information regarding stocking was obtained from communications with Henry J. Vondett, Michigan Department of Conservation. 8' nl I I‘ ll" Table 15 Record of trout plantings for McCoy Creek l9h8-1962 92 Date Brook trout Size Rainbow trout Size 191!-8 hOO Legal -- -- 19h9 700 Legal 200 Legal 1950 900 Legal -- _- 1951 1,200 Legal 100 Legal 1952 1,300 Legal 100 Legal 1953 900 Legal hOO Legal l95h 1,200 Legal 100 Legal 1955 1,200 Legal 100 Legal 1956 600 Legal -- -- 1957 600 Legal -- -- 1958 1,200 Legal -- -- 1959 1,200 Legal -- -- 20,000 Fry 1960 1,200 Legal -- -- 1961 600 Legal -- _- 1962 600 Legal -- -- (i'llunr 93 To provide some data regarding a general index to fishing pressure, species and size of the fish harvested, a fisherman questionnaire form.was prepared. A sample form is duplicated below. Date n_:“_ Time spent fishing Number of legal size trout caught Size of trout: 7-10 in. ll-lh in. 15-25 in. (check one or more) Number of sub-legal size trout caught and released Kinds of fish: Brook trout Brown trout (Check one Sucker (Chub Other or more) Four easily accessible locations were posted with streamrside signs soliciting information regarding fishing results. Questionnaire forms were located in metal con- tainers along with a pencil, at each station. It was requested that upon termination of fishing, a form be completed and deposited in the box provided (Figure 19). Due to limitations of time, no attempt was made to contact fishermen personally for interviews or to check on the percentage of fishermen actually complying with the request. The general prankster who completes questionnaires fellaciously introduces errors. Forms obviously of this type were discarded. Data from the station in Buchanan were primarily of this type. flu Plate XI Fig. 19. Set-up for soliciting information from fishermen. 95 Undoubtedly, some data were inaccurate due to in- adequate knowledge of fish identification. Some data were "missed" due to an absence of some piece of equipment from the-station. All stations were checked frequently to reduce occurrence of this situation. It was almost impossible to keep the Buchanan station con- stantly supplied with materials. After the second week of the trout season, no more attempts were made to keep this station "active." Unfortunately the wooden box at the Bakertown-Buchanan bridge site was destroyed by vandalism. .All data were lost. Results.--The author resided withhobservation distance from.the M-6O station and drove by the Bakertown-Buchanan bridge15 daily. Comparing the observed amount of fishing activity with the total number of returned questionnaires, it appears that many fishermen did not comply with posted requests. or the 39 usable forms returned, 95 Percent of the 36 fishermen indicated they had some success. It is highly doubtful if this high success ratio can be pro- jected to all McCoy Creek fishermen. Needham.(19h7) re- ports that over 80 percent of the fish taken in any given watershed are taken by less than twenty-five percent of the anglers. This would tend to corroborate the author's personal observations of high use, but few returns. 15 By visual observation, the most heavily used fishing site. 96 Shetter (1951) stated that streams with concen- trations of large adult trout will receive the greater part of their angling pressure and the greater part of the season's total catch will be removed during the first four weeks of the trout season. Morofsky gt_gl. (l9h9) found in a southern Michigan study that lhh of the total 180 brook trout reportedly taken for the season, were reportedly taken on the opening day. This evidence tends to explain the marked decrease of fishermen visually noted using the stream after the second week. Fishermen were visually observed to concentrate at the two fish release sites. Cooper (1952) noted similar results stating that planted areas attracted about three tbmes as much fishing as unplanted sections. Analyzing the data obtained and summarized in Table 16 a total of 82 legal sized trout were reported to have been caught. Ninety-four and five-tenths percent of these were brook trout and 6 percent were brown trout. Eighty-nine and five-tenths percent of the brook trout were of the size planted (7-10 inches). The next size group represented 10.5% of the reported catch. This class could be survivors of last year's plant, extra large fish of the 1962 plant, or another age class from natural reproduction. Since no brown trout had been stocked in recent years, those of legal size reportedly caught must have been the result of natural spawning. Table 16 Summarization of catch data from fishermen questionnaires 97 e do uoaoemn you sense Hope» he uncomom m H n as m on «H oHeaoa n H m o: : om 0H eweHaa room oaahhsm H H on : om m can: cohoapnen epHu owoann nsoemosmnnkouaemem un nu nu m uu m some canoam mesenosm condo .nH .nH moped .GH .nH HeMoH ,II o o n u Homes ,anHH pans unsm noHu-em cream nun mooam unwise popaomea psonmiMo aopfisz 98 Unfortunately, the sub-legal size trout were not listed as to species. Other fish listed as caught were chubs, suckers, and mud pickerel mentioned in the order of des- cending frequency. Discussion.--Under "normal" population conditions, it could be expected that bait fishermen would be catching trout other than just "keepers." Shetter and Leonard (19h2) published results from a brook trout population study from a limited area indicatinglé; 2.3 percent were 7 inches or more in length, 22 percent between h inches and 6 7/8 inches; and 75.7 percent were less than h inches. Schuck's (19h5) findings for brown trout showed approximately 56 percent of the fish in age class 0; 23 percent for I; 11 percent for II; 6 percent for III; 2 percent for IV and 1 percent for V. This information suggests that data ob— tained for McCoy Creek's trout pepulation shows it to be primarily an artificial one. Assuming the sub-legal length trout reported 1232 trout and not chubs, only 1h percent of the total catch was sub-legal, while 86 percent was legal. It must also be considered that the hook size would tend to eliminate catching some of the smaller fish.17 Generally 16 The authors stated that for their study it was impos- sible to predict accurately the ages of the various elements of the brook trout population from their total length. 17 Problems of Trout Management, Michigan Department of Conservation, Fish Division Pmmphlet No. 13, l95h. 99 though, it appears that the native population is materially augmented by the planted trout. Assuming all sub-legal trout were brown trout (they evidently spawn in the creek), they would represent 70 percent of the total brown trout catch reported. Sizes 7-10 inches, 2h percent; and sizes ll-lh inches, 6 percent. This seems to be more in line with data presented from.the literature. On this basis, the McCoy Creek native popu- lation might be considered following a somewhat general and standard pattern, but one of very low productivity. Electrofishing Thepurpose of this phase of study was to obtain samples of the streamfls larger-sized resident fish popu- lation, arriving at some indication of their abundance. Estimates of the total population by species were beyond the scope of this project. Six areas were selected for sampling: one each in Section II through V; two areas for Section VI, and the largest and deepest pool in the stream (also in Section VI). The areas in each section were chosen for their accessibility as well as being representative of each section.18 The work was carried out in December, 1961. A pair of hand-held electrodes each connected to 100 feet of insulated wire leading from the generator, 4 18 The pool being an exception. 100 served as the "shockers." A portable 500 w, 110 v, 60 cycle AC generator provided the power. By locating the generator midway in the section to be worked, it was possible to shock a total stretch 200 feet long. A blocking not was anchored across the stream 200 feet upstream from.the in- tended starting point. The 100 foot leads were stretched out on the bank prior to entering the stream to reduce the chance of scaring the query. Two men worked in the electrodes upstream.towards the not. They strived to cover the entire stream without al- lowing fish to swim out and around the effective "charged" area. The distance between the electrodes varied. Where pools were encountered, a narrower but more intense electric field was desired. In clear, clean-edged, shallow stretches, a wide field sufficed. One man followed, netting the fish. Very few fish that were observed to have been stunned, were lost. No estimate is available of how many unobserved stunned fish were missed. No attempt was made to not all the smaller fish, but a note was made regarding their observed frequency of occurrence. Approximately 1,250 feet of stream.were shocked. This represented about a 2 percent sample of the total stream's length. A summary of the results is tabulated in Table 17. Results and Discussion.--Electrofishing using an AC generator (500 w, 110 v capacity) has proven valuable for Table 17 Results of electrofishing--by sections. Ratio of stream.worked to total area of stream comprising each section 101 Hanmnsnv H H H m mH om eonao opunEmonpmaa been oom .oweHan noooot n.3eaond no.8uenumaaoo Hoom pmth Bonk aH> NH mm Eeonnnmn noon oom .nH noHeeem mm on Buchanan been oom .omoHae unoEeo u.oeom hnopofioo_sohm >H mm on m QOHaepu no Esoaamosoo neon om HHH MN m: pooh oom Buchanan :oHpeaa .emoHan n.08onm moons Beam HH oomeomo obon on none none eczema HegemoHQ steam moose one: seam: cox nuwm .mu nhqaom mmwaowram mononm m hobo meHmemm nmwh nooawoo .qfie» AH< .QEop Hepsz AN easth oeuv ooMoonn coed noHpoen aaohpm 102 .Hopaeo .AeusonnHav GHQHdon .eoom .eoneaHSooo no honesoeah oebhonpo ho neono ca mono: mean hoHHeEm * hobo canopy HHH> aGOHQEmm 902 .NH .HH> 0H one mH nu nu nu uu uu NH mm uaoHueum nee?» non Hoom owaaq 0H> Buchanan been OON N N NH pm .oH noHaepe Bonn antennas pooh o: nH> oomnomo eooameo e>mn on same page hexane Hoaomoan .maep .gao» AN shaman eemv :oHuoou steam moose one: epan on: aH4 Home: oemoone seam Eeonam anmn .oz uhaaom sumomwnH mononH m nopo moHoomm anm 103 fish capture. Its efficiency is variable, being more successful in turning up the larger fish (Shetter gt_gl., l9h6; Cooper, 1951; Pratt, 1951; Benson g£_gl., 1959). Larimore (1961) studying a warm.water stream in Illinois found its efficiency to vary with the behavior, habitat and morphological characteristics of each fish species. Pratt's (1951) work on two southern Michigan trout streams with AC and DC generators also indicated species variation in relation to ease of capture. Trout listed in apparent order of difficulty of capture were brown, rainbow, and brook trout, the brook trout showing the best capture rate--lOO percent.19 Shetter g§_gl. (l9h6) working with brook trout in the Pigeon River obtained excellent results; efficiency of capture averaged 96.7 percent, varying with the size of the fish. Benson g£_gl. (1959) found no sig- nificant difference in recovery rates between brown and rainbow trout. It might be conjectured that warier browns moved upstream.out of the area in greater numbers, since no blocking not was used. Cooper (1951) publishing on two year's data regarding brook and brown trout shocking re- covery, indicates-no significant difference between species in the rate of recovery. The collector's ability to see and collect fish, the weather and water conditions can vary the computed 19 Very small sample. 10h effectiveness of the shocker and data obtained (Shetter g§_gl,, 195k; Larrimore, 1961). Considering the reported efficiency the most signi- ficant results were negative-~no trout! Since 600 legal brook trout were planted during 1961 and they may be some- what easier to capture, it was assumed that some would have been taken. The absence of trout from the catch suggests several things: 1) a high percentage of fish planted are caught by fishermen soon after planting. Lemmien gt_gl. (1957) published a recovery rate of 66 percent for rainbows in a southern Michigan stream. Cooper (1951) showed a 6k percent recovery rate for brook trout and 67 percent for rainbow trout in the Pigeon River. Cooper (1953) also found a 59.5 percent return in the same stream for 1951. He stated that rainbow trout planted in Michigan during the angling season at accessible points should give a return in excess of 67 percent. This seemingly would be applicable in McCoy Creek where the initial angling pres- sure is heavy and the plants are made just prior to and during the open season. However some recovery rates in southern Michigan streams (Shetter, 19h?) showed much lower percentages; 17 percent for rainbow trout and 12 percent for brown trout. 2) A high rate of mortality for planted fish has been thoroughly demonstrated in the literature. It seems reasonable to believe that assuming 30-h0 percent20 20 Based on Cooper's estimated 67 percent return. 105 of the trout planted escaped the fisherman, a large number of these would have succumbed by December. With only a 2 percent sample shocked, the probability of taking any of these seems slight. Another explanation may be: 3) An absence of native brook trout and brown trout in the sections sampled. Section V was very difficult to sample effectively. This section, however, appears to contain certain characteristics of a more favorable trout environment. h) The trout moving out of the colder waters into warmer sections. "Typical" samples were shocked however, in upstream.warmer;water sections without results. 5) Still another explanation for not recovering any trout is an inherent sampling error, i.e., although trout were present in the large sections sampled, none were in the units shocked. Mud pickerel (E325 americanus), were recovered in all sections, appearing to be the most universally distributed and abundant fish in the stream. This concurred with the opinion formulated by the author from fish observed during survey trips. Other fish listed in order of frequency of capture were: Northern Creek chub (Semotilus atromaculatus), white sucker (Catostomus commersonnii) and horny-head chub (Nocomis biguttatus). One sunfish (Lepomis spp) was captured, but lost. 106 Dace and sculpin (Cottus bairdii) were the most fre- quently observed smaller fish associates. It appears that the trout population in McCoy Creek consists largely of planted brook trout. After these trout are caught or succumb naturally, the creek's trout popu- lation will be very low. What trout that may exist might be located in the more inaccessible regions of the stream. In general, the data suggest an overall fish popu- lation composed of few species and low numbers. Minnow Seining Various workers have described the significance of the presence or absence of smaller non-game fish. These forage fish assist in designating the ecological en- vironment of a stream. They also serve as forage for the larger, carnivorous fish. Conversely they compete with the same carnivorous game fish for the streamis food organisms. Because of this competition the best brook trout streams would be those which contain only trout and no forage fish. Brook trout make very satisfactory growth on bottom organisms without foraging on smaller non-game fish. Certain cold water associates such as Northern Mudder and Miller's Thumb are thought to be destructive to trout fry (Murray, 1938). Collecting took place on June 13 and 1h, 1962. The seining was done at certain previously located sampling 107 stations which represented visually different habitats (see Figure 2). A six foot by four foot Common Sense minnow seine was used having a 0.125 inch square mesh. Two men, each handling one of the seine's two poles, made 3 to h sweeps approximately 6 feet long along the streamis bottom. All individual sweep samples were collected and placed in sep- arate jars. Later, living material was transferred to jars containing a 5 percent formalin (with borax) solution. Identification took place at a later date. ‘ I Data recorded at each sampling site included bottom type; air and water temperatures; water depth; number of sweeps at each location; approthate velocity (using velocity data for stations obtained at time of November survey). Table 18 summarizes the data collected. Results and Discussion.--Data of Table 18 indicates the app. in order of abundance to be: Number *Rhyinicthys attratulus g. ............... 37 igyprinidae larvae ....................... 23 Catastomidae larvae ..................... l6 _Etheostoma caeruleum..................... ll trimephales notetus g, ................... 10 Cottus bairdii g. 6 GOttidae larva. ......OOOOOOOOOOOOOOO.... 2 108 Number Etheostoma nigrum g, .................... 3 Etheostoma blennoides ................... 1 *Etheostoma microperca ................... l Semotilus atromaculatus g. .............. 1 egggg_americanus 1, ...................... l Summarizing from.Troutman (1957) all of the species in this list (except those marked with an asterisk) typi- cally occur in streams with a steady flow, medium gradient and clear, unpolluted water having sand and mixed gravel bottoms. Pimephales is reported to tolerate a wide variety of silt bottomed degraded habitats. Etheostoma microperca is more typically a species of low gradient streams with silt bottoms. Egg§_americanus is characteristically found in clear, densely vegetated streams whose bottoms are composed chiefly of organic debris. This latter species because of its size and ability to elude the seine probably represents an accidental capture. Figure 16 portrays the relative productiveness of the seining. As the graph indicates, there is a general correlation between the number of fish captured per sweep, the percentage of seining sweeps producing fish and the type of substrate. Since the stream is composed predom- inately of sands and silts, the data suggest environments not conducive to high potential populations of forage fish. Table 18 Minnow seine data Obtained June 13 and 1h, 1962 I- . A . * a. 109 anaanewwl gpmoofim H .e usaspanpuu awake Hopenm .HIIIfiIIn £23 am A S .3: $ 1: m} 8.3.3: a. .>Immmdmflwoam awake - Noam H ma .cox nu nu m\N onam 0 30am - wanton 3 . .3: me E. <0 u: .656 m .p ammnmem roan - m no 3:38 a .. .32 me E. a} noun 533% waive; ma :on mo ma. m\o pduuenam : é -. in no esopmoospm H ma 30am mo ms H\H paanaonmw aaonpnms .mmflmmmwm saopmdonpm : .a azaspapumm phase Hebahm mwnpnflfinm P B so: no me Rm 3E m BF wnanuoz 0H koam mo :5 «\o paueuonam N .o.wmflmmmmmmmz 30am shunnemnhnm w mum .mo: mucus H HH snag mean .na Npao nee ec henaun nhuhn Hagen uoHob .mfio» .QEep Hapop .oz newcomu hope: .Noaamd hopes. pa¢ \noosa epshaepflm uoapepm seapoem one .on o>aaosc nopm 110 abHsneho nacho «sapwoonpm m mH .. mm H» H\H nope; 5H memmonner H awaopmoonnw H L»: asymH: thzm Hopehw asawuoospm H mm .eoz mm Hp uxw oaam-eqam NH WWI oapneH aqum m oaeaaaseho m mH .eo: am ma H\H pHau-eaam :H xfisoHsnoao «Bowmoomww N m .n HHmeHam amaze Ho>asm :H-MH moppoo H oH .co: pm me o\: oqau-eoz noospom ospaaH 3oHu mHaHH osmHsoamomam.m :H .eox om! we :xH enam soozsom > ombnaH onm oneacasnho e m .eoz om ea :xH esam HH wMImumpmm moHanneSHm OH s>naH puHsn Ho>enm eoaeapcoeanp H :H .00: we as m\m onau-e:am OH .9 Hmmnmmm asp»OU.H H BuHsaeuo ummmmmmmmMM.H aoHpupm .: asmmHn pqun Hobshw .anem «soumoonpm H :H .eo: mm as mx: usam-mnmm o >H anu nuHm .:H tho scammec Mensa: nhuho Apnea noHo> .980» .9209 Hapop .oz ueHoemm have: .Konnnd nopuz 9H< \noozn epshpunzm quuapm :oHpoem one .o: o>Huosc loam 111 m eabpmH puHsm 0:00 MNINN 000Hmthho mH :Nan .002 m0 we :\H nmxoom noosvom 00>n0H 000H80p00pao H H 00>90H ppHam Hobapw omcappmb m 0H .00: me 00 me osHmnmasm mm [NH .2 Esan: «Eopmoospm H .n Hmmammm menace H HH :OHm-co: me on m\m pHHm-ecmm am a 0:00 EH 00000980 quspoz oH puwmm me 00 me msooe .mH mm HHH> .n HHOhHmn axooh mappoo H 0am : .0 msHsW0auvm Hoganm HNnom msmwmothnm o ownaq aeozuom HH> owphmH H0>0nm1 oaeHsopmapao 0 mm pqum mo 05 m\H quu-0oz 0H .0 m5H590Auua manpnoHcmmm 0H .0 mspmHSowEonwm a: 080m v H . HHHA H 0» mm o 00 n HHO Han H 0 OH H A0.vn V mopuoo H m pmHzm m0 lmm ooHu-eoz.‘ swamped: 1m> SmHh anm .QH huHo 000mw00 M09852 nmmmb na900 uOH0> .QE0p .msop Have» . .oz noHoomm hens: .Monmm< M0903 9H4 \90030 onehpansm noHpapm GOHpoom 0cm .oz 0>Hnos0 loam H2. PERCENT OF sax—£95 Paooudma CiiaH L $9 8 3 5% if: 9: KS 3 6936396 Rim/£1. SaMD- SILT ‘ I L0. 0. m. 0. l"? (\1 N .__-o H Bap. GRGPH-NumBER OF FISH PER awe-E? meuum-FWL @KRVEL ‘iflND‘FNL eaavh. 5WD E ig are 16 eininv DrOTUUt1V1ty to substrate. 4‘.___—— 113 Hallum.(l958) and Cooper (1952) list some species typically associated with trout in streams such as Rhynichthys atratulus, Semotilus atromaoulatus, Cata- stomidae and Cottus bairdii. All of these occur in McCoy Creek. Some Percidae (Ethestoma spp.) are also evident in the stream's gravel riffle area. As previously stated it is generally agreed that high quality trout stream habitat usually has a high trout/forage fish ratio. Invasion by Per-cidae has been stated (Gutsell, 1929) as an indication of declining trout habitat. McCoy Creek has a low trout/forage fish ratio and Percidae inhabiting the gravel riffles. It is therefore postulated that this trout/forage fish ratio suggests a marginal trout stream environment. Those trout that do exist have to face considerable competition in the various stream habitats, i.e., water cress beds--fingerling pre- dation by Esox; food competition in gravel riffles by Ethestoma spp., and in sand, silt and gravel bottoms with the other array of fish. Possibly with environmental improvement favoring higher trout populations, their competing fish might become forage for an increasing trout population and therefore become more limited in their distribution and abundance. Shetter g£_al. (l9h6) noted that after stream improvement work had been in part completed on the river, llu brook trout and sculpins increased in numbers; the trout had a corresponding increase in weight and rate of sur- vival. The associated minnow population failed to in- crease in weight per fish and showed only a slight increase in numbers. Ground water and Spawning A study by Benson (1953) indicated that ground water seepage influenced trout populations in the Pigeon River by controlling the location of brook and brown trout redds. Population estimates indicated that trout abundance was correlated with the number of redds per area. Recent investigations have indicated that for salmon spawn to hatch (Wickett, 195h) and the embryos to develop 2 22 1960; Coble, successfully (Shumway, 1 1960; Silver, 1961) certain stream bed conditions regarding oxygen and velocity of ground water percolation must be met. These conditions seem even more important in the case of brook trout than brown trout. Various workers (Pollard, 1955; Terhune, 1958; Gangmark, 1958) were prompted to develop apparatus and techniques to measure the velocity and oxygen content of the stream bed. Although these techniques provide more 21 Unpublished H.S. thesis. 22 Ibid. [I I [ll-I. ll. 115 refined data, it was not within the scope of this survey to utilize these methods. Using the general survey methods of Benson (1953). e.g., winter water temperature, surface ice cover, summer water temperature, and gravel substrata temperature, areas with ground water seepage were located. Winter and summer water temperatures have been dis- cussed more fully in an earlier section of this disser- tation. In general, only the main feeder stream defi- nitely showed evidence from seasonal temperatures, of sig- nificant amounts of ground water influence. However, because the substrate in this section seemed unsuitable for brown and possibly brook trout spawning, no further investigations were conducted in this section. Most of the stream.was observed frequently through- out the winter months. Dates of stream.ice formation and its coverage was noted and are depicted in Figures 9 through 1h. Only on the coldest mornings were there evidences of streamredge ice upstream from station 12. The ice cover formed at places of reduced water current (cress beds) and around protruding stream brush, bushes and logs. Below station 12 stream—edge ice was more noticeable. During the coldest periods Section V had an ice-free channel (2-3 feet wide) but an ice layer frozen to the interlacing branches on the water's surface. Section VI 116 had an ice-free channel 2-3 feet wide. The ice was cov- ered with 8-10 inches of snow except during parts of February. A few isolated areas in this section contained no ice and were noted for future inspection. Section VII retained a 2-3 foot open channel with conditions similar to those in Section VI--but there were no areas of "open water." Section VIII was solidly frozen for most of its length during the most of January and February. Section IX had some edge ice present. Industrial and sewage effluents discharged into the stream.epparently warmed the water enough to prevent ice formation except for short periods of time. Extent of ice cover did provide general information about where not to look for ground water seepage. Since all gravel areas (potential spawning areas) had been mapped, a check of the substrata temperatures in these locations was undertaken. Special attention was given to those stream portions which had little or no ice cover during the winter. A battery operated thermometer on which were con- nected 2 wire electrodes was used to take substrata tem- peratures. One electrode was attached to a sharp stick, then probed 2-3 inches into the substrate; the resulting temperature was recorded. The second electrode was kept in the stream to record general water temperature compared 117 with that of the substratum. Tabulation of the survey's results are given in Table 19. Of the gravel beds present in Section III, only one stretch approximately 75 feet long, downstream.from station 7 (see Figure 2) was there ground water seepage through the substrata. The soil temperatures here were 50° compared to 660 stream.temperature. Many stream bank water seepages were noted, but these seeps were flowing directly into the stream's surface water and not entering by way of the substratum, Three substrata water seepage areas were located in Section VI between stations 18 and 19. All three were in the ice-free areas noted during the winter survey and located in the south banks and substrata. These seeps varied in bottom surface area coverage from 120 square feet to 350 square feet. All seeps percolated through gravel of various sizes intermixed with varying amounts of sand. The water depth over the seepages varied between S to 12 inches of medium surface velocity. The substrate in the seepage beds was noticeably'more loosely packed compared to other gravel beds having no ground water activity. Table 19 summarizes the morphometry of the ground water areas located. According to previously mentioned investigations, seepage through the stream.eubstratum.is a very important ecological factor affecting trout populations. Apparently Table 19 Locations of ground water areas graphically located in Figure 3 118 QOHQHHHHOQ Hddhpmfiflfld OS“ Oww3mm ho. UOfiOOHHGIIOOuAGOflO D02 NH hobasm :09:H3 w:HHS0 thpro0 :0903 w:Hnmm no 000:00H>0 o:un00&oono poz HHH> hebaom :0»:H3 w:H::0 th>Hpo0 nova: w:H::0 no 000:00H>0 o:un00xo0:o poz HH> 0:00 :sz 00HHE .pu.du mom mm me Hopesw NH-m 00H: sue . eaHe s.eoz meoH mm-omxm 0:00 :pH3 eoxHa .pe.eu 0H: mm me Hopesw OH-m 00H: oH-m onHu s.eoz waoH om-ozflm 0:00 0Eo0 .ph.vn omH mobaso mm me Hopeam OH-m 00H: m-: . some sumo» so HH0 .mH 0wa0H m:oH omnomaH 0:0 NH 0:0Hp0un :00300m H> :pmo0 nep03 090Hamohmm0 0:0 000p Hebuaw no xe0H o» 0:0 00xoone poz > 00p000H o:oz >H endeHE .au.00 0mm 00:» EH0 cm 00 0:00 NHuOH 00H3 mum 00:3o0 empaH soak 500:9095 H0>0nw.00£ w:oH mwnOQ hH0p0H0onEH .x:0n 9000 um 0:0 w 0:0Hu0pm :003H0m HHH 00xoono po:nu00on H0>0aw 02 HH 0030030 ao:nnm0on H0>0aw oz H 000aw00 .:H .9509 .980» Espohpmnsm :pm00 0w0aeboo Ho 0&09000 he :oHp0ooq :oHpoem 090apm nope: :0p03 pooh eh0zdw Imam 119 from earlier years' fishing, many more trout were caught in McCoy Creek, including those of sub-legal size. If the stream.former1y produced a native trout fisheries a decline in suitable spawning areas may be a major factor in a decreased trout population. Reduced spawning areas may have been a result of a heavy build-up of additional substrate from erosion etc., a decrease in the ground water seepage, and/or a combination of both. The problem of erosion and siltation in the stream.has already been dis- cussed. To fully investigate the second hypothesis would require investigations beyond the scope of this project. However, we can examine some ground water facts and local records and opinions. Spring fed streams receive their water supply pri- marily from the ground water zone known as the zone of saturation (Ackerman gt_gl., 1955). This zone is at the surface level in some lakes, ponds and swamps acting as ground water storage units, feeding streams continually. It is "recharged" from precipitation percolating into the ground. These interactions are part of the hydrologic cycle. The rate of seepage into the streams is determined in part by the hydrostatic pressure (pressure caused by the weight of water above). With proper geological con- ditions existing, sufficient pressure occasionally occurs to cause the water to rise in a well above the land 120 surface creating a flowing or artesian well (Ackerman, 1955). Elementary physics indicates that this pressure which feeds water to wells and ground seepages or to tur- bines at a dam.site can be reduced, if the stored water (zone of saturation in the case of wells and seeps) is exploited faster than it is replaced. How much do we know about water conservation and hydrology? Harrold (l9Sh) states that ". . . our stock- pile of data for meeting water conservation planning needs is distressingly inadequate." Thomas' (1955) views re- garding this lack of information are ". . . that hydrolo- gists can see no farther underground than other people." Information regarding spring activity, well flow, ' stream flow, and general water conditions were solicited from the aforementioned local residents. Of those who felt qualified to answer, all mentioned a reduction in the total volume of stream flow. Many specific incidents were cited, e.g., present stream level (already discussed in an earlier section); reduced frequency of spring floods (3, h); disappearance of swimming holes (11). Former flowing springs in close proximity to the creek have ceased to exist (1, 3, h, 9). These springs according to in- formation received, had a very strong, large flow of water during all seasons. Much of the now ditched muck land was covered by water, cattails and sedge. Now the same area 121 supports a stand of cherry trees, grey dogwood and some crab apple trees (2). Results and Discussion.--Where has the water gone? Examination of the population statistics for the town of 23 shows a 36 percent increase in population from Buchanan 1930 to 1960. Added water demands come from an enlarged Clark Equipment factory and the addition of several "new" factories. On the McCoy Creek watershed in Buchanan” Township, an 80 percent increase in home numbersah has occurred since 1930; an approximate 11h percent increase has occurred in Bertrand Township. Two local well drillers (12, 13) who have been con- nected with the business for 15 to 20 years provided some additional information which is summarized below. 1) Spring water activity has decreased. 2) Wells in close proximity to McCoy Creek are still supplying water at 25-30 feet. 3) Hells along the north part of the watershed are about 60 feet deep and fairly stable. h) Periodically they are called to redrill old drilled wells because they have gone dry. 23 City water is pumped from deep wells near the stream. 2h Figures determined by comparing the number of houses indicated on a 1927 0.3. Geological Survey map with those now present on a 1960 aerial photograph. Results expressed in percent. 122 5) It was their opinion that very few hand-dug wells were still producing. Most of them have had to be drilled deeper. Undoubtedly, increased tiling of farm lands_adjacent to the creek has added to the reduction of hydrostatic pressure as has ditching activities. From.the preceding data it appears that the ground water supplies in the vicinity of the creek, seem adequate for human consumption. The data suggest however, that hydrostatic pressure in the watershed has been reduced. This pressure (or head) has been stated by Pollard,(l955), to be very important in determining the velocity of the 6 ground water seepage, along with permeability of the gravel. Reduction in the stream's substrate permeability has already been demonstrated to be a logical aftermath of the erosionésiltation of former and recent years. Reduction in water head could also account for lack of pressure to force ground water through the streamis sub- strata. This in turn might explain an increase in stream temperature, a reduced stream depth, and a decrease in suitable trout spawning areas. Limiting Factors The foregoing data presented and their discussions suggest certain seemingly outstanding deficiencies in the stream's ecology. l) A shortage of adequate spawning 12} areas. 2) Inadequate production of bottom fauna deter- mined from a food scale rating. 3) Low density of ade- quate adult trout shelter. Pools present are generally small in surface area and only sporadically distributed. Suggestions for Management Arranged in the suggested order of completion. Environmental Improvement Headwater Hanggement.--It should be quite obvious to the reader that little can be accomplished to improve McCoy Creek for trout unless the lands in the headwater area are stabilized. Of prime concern are the bottom lands (Sections 16 and 19) from which much spring water emanates--and is the major source of siltation. The fol- lowing discussion will concentrate on this area. The author suggests several possibilities: 1) Through joint cooperation between the State Department of Agriculture and one of the State Conser- vation agencies, these bottom lands might be put into a soil bank trust or lease predicated on a long term.man- agement basis. Only after reaching a mutual agreement between the land owner, department, and the administering conservation agency, could these lands be put back into production. This should insure future safeguards against a recreation of the siltation problem once costly imp provement measures are completed to "restore" the stream. 12h 2) Elimination of the open ditches by installation of a network of tile drains. According to the local Soil Conservation agent, tiling would provide a more effective drainage system, discharging only relatively clear, cold water onto the stream. To accomplish this, some ar- rangement might be worked out through one of the conser- vation agencies, department, and local landowners. 3) A multi-purpose recreation area might be de- veloped. The area would have to be purchased or leased on a long-term.basis to assure proper management. The area is ideally suited for the establishment of a managed water fowl marsh. One water control structure would be needed near Buffalo Read. Only a limited amount of diking would be necessary because of the nearly ideal surrounding terrain. This area, administered by the Conservation Department could provide a fine water fowl nesting area. If desired, regulated (or open-to-the—public) shooting would create excellent local interest and recreational opportunities. This arrangement would undoubtedly raise the water temperature of the stream. This increase, due to a read- justment of the streamis heat budget, could be approxi- mated by computation. This increase could be offset, in part, by tapping the spring water under the large mounds scattered through the marsh. The water has varying amounts of vertical pressure which could conceivably be capped and controlled. 125 Literature has already been discussed concerning optimum and maximum.water temperatures for brown trout. Near optimum.water temperatures for brown trout could be maintained by installation of stream improvement devices. The many cress beds now present are valuable for trout fingerling cover. Shelter for adult trout is greatly needed. Properly located stream.1mprovement struc- tures should increase the number of pools, undercut banks, etc. Increasing_Production of Bottom Fauna.--l) The pre- ponderance of silts and sands forming the substrata ap- pears to offer an inhospitable environment for most aquatic fauna. It is the author's opinion that to assure water velocity of sufficient swiftness to maintain an unsilted bottom, the total stream width in Sections II, III, and IV should be reduced 3-h feet and 5 feet respectively. 2) Addition of rocks on the scoured bottom could create riffle areas and in general, a more suitable habitat for bottom fauna. Gravel size similar to that used along railroad grades should prove adequate.25 Creation of Spawning Areas.-—Various workers (Needham, 1961; Stuart, 1953; Webster, 1962) have demonstrated the possibility of creating artificially attractive trout spawning situations. Further field application of these 25 Figure 12, results of sample #9. 126 methods may provide the necessary additional spawning grounds in McCoy Creek which now appear to be very limited in their distribution. Investigations by the author using field tiling systems to simulate ground water upwelling through the substrata are being contemplated. Trout Populations I Recommendations for management of the game fish popu- lations would vary with the methods used to stabilize the watershed. If the present water temperature is maintained, or decreased through various measures outlined, a native brook trout fisheries along with improved brown trout con- ditions, might be developed. If the water temperature permitted only brown trout inhabitation, pro-season and early season stocking with brook trout would provide fast but short-lived sport for the first two weeks. The brown trout would provide a good native population testing the skill of the trout fisherman and providing him with a good residual population which contained older and larger fish. An alternate to the trout management program.would be a serious evaluation of the stream.for northern small mouth bass (Micropterus dolomieu) occupancy. SUMMARY Investigation of an eleven mile long stream.(McCoy Creek) in southwestern Michigan was prompted by the scar- city of trout stream information available for this portion of the state. The stream's past history was compiled with the aid of two public agencies and eleven selected local residents. Historical information indicates major dredging occurred in 1897; other significant bridge construction and dredging work was carried out in the mid-thirties and early forties. All survey and map work was completed by wading or canoeing in the stream, and walking the bank. The stream was divided into nine sections and morphometric data were compiled for each section. Little attention was given the last section due to the water's constant exposure to industrial and sewage wastes. The substrata were composed primarily of silt and a sand-silt combination. Other types in their order of abundance: sand-fine gravel, sand, medium gravel, and small rocks. Medium.gravel and rock harbored the most bottom fauna; silt and sand-silts were the least pro- ductive. Water cress and Anacharais yielded good samples of fauna-~primarily crustaceans. Vallisneria and Entamggetgn pusillus harbored a rich fauna in the summer 12? 128 only. Tendipedidae, Cylloepus and Hydropsyche were the most common insect types found in the samples. This stream appears to be low in aquatic fauna production. Most of the stream was composed of moderately slow water (under one foot per second). The fastest water (over two feet per second) was in the lower one-sixth of the stream. The gradient averaged 1.2 feet per 1,000 feet for upper 8h percent of the stream's course; 59 feet per 1,000 feet for the lower 16 percent. Water shading by bank shrubs was fairly good, ranging from very little (Section IV) to almost 100 percent cover- age (Section V). Shading apparently had some effect on water temperature during certain warmer months. Water cress was fairly well distributed and spor- adically plentiful. Most other aquatic plants varied in abundance with the seasons. Very few logs, undercut banks or stream debris, were present. The predominant water type for the stream was flats, and much less common runs and riffles. The streamrpool area ratio was very poor. Results from fishermen's questionnaires indicated the catch to consist primarily of brook trout, 7-10 inches long, and some slightly larger. A few 7-1h inch brown trout were reported. A small number of sub-legal sized 129 trout were reportedly caught. The native trout population appears to be composed of brown trout with a low popu- lation density. A sampling of 2 percent of the stream by electro- fishing returned no trout. Mud pickerel was the most common fish with a few horny-head chubs and white suckers recovered. Seining results indicate a correlation between forage fish abundance and distribution with substrate type. The trout/forage fish ratio appears to indicate an unfavorable competition situation for the trout. The species of for- age fish inhabiting the stream suggests it to be one of marginal value for trout. Crayfish appeared to be quite abundant in the sand and gravel substrata.. Only h substratum seepage areas were located. Their importance to trout spawning was discussed. A general reduction in hydrostatic pressure in the creek's water- shed seems to have occurred over the years. This re- duction was evaluated in terms of increases in rural and urban populations. The major limiting factors appeared to be: 1) a shortage of adequate spawning facilities. 2) Low aquatic fauna production. 3) Lack of shelter for adult trout. ‘ ‘ r LIST OF INFORMANTS Length of—_9 Fished stream Names of Informants** residence consistently yrs. Yes* No 1. Coleman, John 50 X 2. Ferguson, Ivan 66 l925-l9h0 Buffalo Road to M-6O 3. Frame, William,Sr. 80 X h. Haas, Walter ‘ 55 X 5. Hanover, Roscoe no 1928-1955 M-60 to city limits 6. Hassinger, Victor 18 7. Kingery, William 20 8. Klasner, Floyd hO 1920-1961 X 1.Entire stream 9. Redden, John 81 2.More recently _ upstream from 10. Reinholdt,Lawrence 25 H-6O X 11. Rohl, Edward h3 1928-1930 1.M-6O to RR grade. 2.Inside city's limits 12. Ferris, Carl Well driller 13. Schutze,Fred J. ‘Well driller * Includes years in which the stream fished most con- sistently and the area of the stream most utilized. All currently residing in area except William.Frame, Sr. who died in January, 1962. 130 LITERATURE CITED Ackermann, William.C., E. A. Coleman, and Harold O. Ogrosky. 1955. From ocean to sky to land to ocean water. The Yearbook of Agriculture, pp. hl-Sl. Supt. of Doc., Wash., D. C. Allen, K. Radway. 1951. The Horokiwi Stream. New Zealand Karine Department Fisheries Bulletin No. 10. Anderson, Richard O. 1959. A modified floatation technique for sorting bottom.fauna smmples. Limnol. and Ocean., Vol. h, lo. 2, pp. 223-225. Anmitage, Kenneth B. 1958. Ecology of the riffle insects of the Firehole River, Wyoming. Ecology, Vol. 39, No. A, Benson, Norman C. , 1953. The importance of ground water to trout popu- lations in the Pigeon River Michigan. Trans. No. Am. Wild. Conf., Vol. 16, pp. 270-281. Benson, Harman 6., Oliver B. Cope, and Ross V. Bulkley. 1959. Fishery management studies on the Madison River system in Yellowstone National Park. U. 8. Fish and Wild. Serv., Special Sc. Rpt., Fish, lo. 307. Boussu, Marvin F. l95h. Relationship between trout population on cover on a small trout stream. Jour. of‘Wildlf. Hgt., VOI. 18, NO. 2, pp. 229-2390 Brasch, John, James McFadden, and Stanley Kmiotek. 1958. The Eastern Brook Trout. Wis. Cons. Dept., Pub. 226, ll.pp. 131 132 Coble, Daniel W. 1961. Influence of water exchange and dissolved oxygen in redds on survival of steelhead trout embr 03. Trans. Am. Fish. Soc., Vol. 90, No. A, pp. 69-u7u. Cooper, Edwin L. 1951. Rate of exploitation of wild eastern brook trout and brown trout populations in the Pigeon River, Otsego County, Michigan. Trans. Am. Fish. Soc., Vol. 81, pp. 22h-23h. Cooper, Edwin L. 1952. Growth of brook trout (Salvelinus fontinalis) and brown trout (Salmo trutta)in the Pigeon River, Otsego County, Michigan. Mich. Acad. of Sci., Arts and Letters, Vol. XXXVIII, pp. 151-162. Cooper, Edwin L. 1953. Returns from plantings of legal-sized brook, brown, and rainbow trout in the Pigeon River, Otsego County, Michigan. Trans. Am. Fish. Soc., Vol. 82, PP. 2 5-280. Cordone, Almo J., and Don W. Kelley 1961. The influence of inorganic sediment on the aquatic life of streams. Calif. Fish and Game, Vol. h7, No. 2, pp. 189-223. Curry, LaVerne L. 195M. Notes on the ecology of the midge fauna (Dip- tera Tendipedidae) of Hunt Creek Montmorency County, Michigan. Ecology 35 (h). Ellis, Robert J., and Howard Gowing 1957. Relationship between food supply and condition of wild brown trout in a Michigan trout stream. Limnol. and Ocean., Vol. II, No. A, pp. 299-308. Ellis, M. M., B. A. Westfall, and Marion D. Ellis l9h8. Determination of water quality. U. 3. Fish & Wildlife Serv., Res. Rept., No._9. 122 pp. 133 Embody, G. c. 1921. Concerning high temperatures and trout. Trans. Am. Fish. Soc., Vol. 51, pp. 58-6u. Frost, W. E. 1939. River Liffey Survey II. The food consumed by the brown trout (Salmo truttg Linn.) in acid and alkaline waters. Proc. Roy. Irish Acad., h5 (Sec. B, No. 7), 139-206. Fry, r. E., Jr. l9h7. Temperature relationships of Salmonoids. Proceed. Nat'l. Comm. Fish Culture, 10th meeting. Appendix D. Gangmark, Harold A., and Richard G. Bakkala 1958. A plastic standpipe for sampling the streambed environment of salmon spawn. Calif. Fish and Game h2:37-39. Goodnight, C. J., and L. S. Whitley 1960. Oligochaetes as indicators of pollution. water and Sewage Wks., 107, 311. Gutsell, James S. 1929. 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Brown 00., Dubuque, Iowa. M21 pp. Lane, A. C. 1907. Summary of the surface geology of Michigan. State Board of Geological Survey, Lansing, Mich., Larimore, R. weldon 1961. Fish population and electrofishing success in a warm-water stream. Jr. of Wildlife Mgt., V01. 25, NO. 1, pp. 1‘12. Lemmien, walter A., Peter I. Tack and Walter F. Morofsky 1957. Results from planting brown trout and rainbow trout in Augusta Creek, Kalamazoo, Michigan. Quart. Bu11., Mich. Agr. Exp. Sta., East Lansing, Vol. no, No. 2, pp. 2&2-2h9. Maciolek, John A., and P. R. Needham 1951. Ecological effects of winter conditions on trout and trout foods in Convict Creek, Cali- fornia, 1951. Trans. Am. Fish. Soc., Vol. 81, PP. 202-217. Morofsky, W. F. l93u. A preliminary survey of the insect fauna of some typical Michi an trout streams. Jr. of Econ. Ento., Vol. 2 , No. 1, pp. 82-86. 135 Morofsky, W. F., P. I. Tack, and W. A. Lemmien 19h9. Recovery of a Southern Michigan trout stream. Quart. Bull., Mich. Agr. Exp. 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Fish and Game, Vol. #7, N00 1, ppe 27-390 136 Pennak, R. W., and E. D. Van Gerper l9h7. Bottom fauna production and the physical nature of the substrata in a Northern Colorado trout stream. Ecology, Vol. 28, No. 1, pp. h2-h8. Pennak, Robert W. 1953. Fresh-water invertebrates of the U. S. The Ronald Press Co., N. Y. 769 pp. Pollard, R. A. 1955. Measuring seepage through salmon spawning gravel. Jr. Fish. Res. Bd., Canada, Vol. 12, NO 0 S , pp 0 706-711.]. 0 Pratt, Virgil S. 1951. A measure of the efficiency of alternating and direct current fish shockers. Trans. Am. Fish. Soc., Vol. 81, pp. 63-68.. Rees, William.H. 1959. Effects of stream.dredgings on young silver salmon and bottom fauna. Fish. Research Papers. ‘Wash. Dept. of Fish., Vol. 2, lo. 2, pp. 53-65. . Schuck,neward A. 19h5. Survival, population density, growth and movement of the wild brown trout in Crystal Creek. Trans. Amer. Fish. Soc., Vol. 73 (l9k3), PP. 209-230. Shetter, David S., and Albert S. Hazzard 19h0. Results from plantings of marked trout of ‘ legal size in streams and lakes of Michi . Trans. Am. Fish. Soc., Vol. 70, pp. hh6-fi68. Shetter, David S., and Justin W. Leonard l9h2. A population study of a limited area in a Michigan trout stream, September, l9h0. Trans. Am. Fish. Soc., Vol. 72, pp. 35-51. 137 Shetter, David S., O. H. Clark, and Albert S. Hazzard 19h6. The effects of deflectors in a section of a Michigan trout stream. Trans. Am. Fish. Soc., V01. 76, pp. $8-278. Shetter, David S. l9u7. Further results from spring and fall plantings of legal-sized hatchery-reared trout in streams and lakes of Michigan. Trans. Amer. Fish. Soc., Vol. 7H, PP. 35-58. Shetter, David S. 1951. Variations in fishing pressure and catch during various portions of the trout season on some Michigan trout waters. Prog. Fish. Cult., pp. 213-218. Shetter, David 5., Marvin J. Whalls,. and O. M. Corbett l95u. The effect of changed angling regulations on a trout population of the AuSable River. Trans. of Nineteenth N. Am. Wild. Conf., Pp e 223-238 e Shumway, D. L. 1960. The influence of water velocity on the devel- opment of salmonid embryos at low oxygen levels. M.S. thesis, Ore. State College, Corvallis. 1&9 pp. Silver, S. J. 1960. The influence of water velocity and dissolved oxygen on the development of salmonid embryos. H.S. thesis, Ore. State College, Corvallis. 50 pp. Smedley, Harold H. 1938. Trout of Michigan. Lansing, Michigan. Stuart, T. A. 1953. Water currents through permeable gravels and their significance to spawning salmonids, etc. Nature, l72:hO7-h08. 138 Taft, A. C., and L. Shapovalov 1935. A biological survey of streams and lakes in the Klamath and Shasta National Forests of California. U. S. Bureau of Fish, mimeo. Tarzwell, Clarence M. 1937. Experimental evidence on the value of trout stream improvements in Michigan. Trans. Amer. Fish. Soc., Vol. 66, pp. 177-187. Tarzwell, C. H. 1938. Factors influencing fish food and fish pro- duction in southwestern streams. Trans. Am. Fish. 800., V01. 67, pp. 21"6-255' Tebo, L. B., Jr. 1955. Effects of siltation, resulting from improper logging on the bottom fauna of a small trout stream in the southern Appalachins. Prog. F1811. cult.) Vela 1, NO. 2, pp. 6h'700 Terhune, L. D. 1958. The mark VI groundwater standpipe for measuring seepage through salmon spawning gravel. Jour. Fish. Res. Bd., Canada, 15:1027-1063. Thomas, Harold E. 1955. Underground sources of our water. The Yearbook of Agr., pp. 62-78. Supt. of Doc., Wash., D.C. Trautman, Hilton B. 1957. The fishes of Ohio. The Ohio State University Press. 683 pp. Veatch, J. L. 1937. Land types and orchard production in Berrien Co., Mich. VonDett, Henry J. Personal communication. 139 Webster, Dwight A. . 1962. Artificial spawning facilities for brook trout, Salvelinus fontinalis. Trans. Am. Fish. Soc. 91 (2), pp. 168317E. Welch, Paul S. l9h8. Limnological methods. The Blakiston Co., Phila. 381 pp. Wickett, W. P. l95h. The oxygen supply to salmon eggs in spawning beds. Jr. Fish. Res. Bd., Canada, Vol. 11, No. 6, pp. 933-953. , e "a Q on ,MJ .f .w-jé 5.6 u he 7‘. I, 9, ‘L’. .1 L. - -id “' 1 Vim HICH iiiiiiiiiiiiIf