Kl APPUCATION OF THE EQUATION OF THE MW GF DIMINISHING {NCREMENT TO SWINE NUTRITION Thesis foe the Dogm of M. S. MRCHIGAN STATE UNIVERSITY V‘erl E. Headiay 1960 APPLIGKPIQI CF THE EWM‘IW W THE CURVE (F DIHIHISHING INCREHENT TO 5m MIG Val E. Hadley MW Sub-uteri to the Cellege of Agriculture Michigan State University of Agriculture and lpplied Science in partial fulfillment of the mural-ate for the degree of MASTER CF SCIENCE Dope-taunt. of. Animal Husbandry 1960 ABSTRACI' Verl E. Headley Two studies were conducted using live weight-feed consumption data from280babypigs {birthtoueaning) cnasyntheticnilkdietandfrcn 1200 animals (weaning to whet) fed a typical ration of fixed ecu-posi- timinanattemttcdetermineifthisdataccnfcrmedtotheequatim of the curve-of diminishing increment and till: Qply all equations derived from it. Averages of live weight and feed consunption were calculated at equalintervalsoftine, thatishdayintervdsforthebabypigsand 2weekintervalsfortheaninalsfronweaningtonarket. The constants of the equation of the curve of diminishing increment were calculated and adjusted by the statistical method of least squares. Its application to the baby pig data on a synthetic milk diet, though reasonable acctn-acy was obtained, was not found to be practical. . since the live weight-feed consumticn relationship of the baby pig over this range (6 weeks) was essentially linear. ‘ bplyingtheequationtoswinedatafromweanixg tomarketwasfound to conform with a high degree of accuracy. Calculated values of emails- tive feed comunption were in agreemnt with 11.8.0. standards. The 1133th efficiency of feed utilisation was equally ids- tica‘l to the N.R.C. standards to 160 pounds live weight. However, from 160 pounds to 200 pounds live weight the N.R.C. values denote a higher efficiency than the calculated values in this stub. Calculated instan- taneous efficiency decreases linearly with increasing live weight. Maintenance requirements as applied in this study and as defined by the equation include all feed consumed exclusive of that which is Verl E. Headley utilised for protein storage (growth). Cumulative maintenance require- nents increase exponentially with increasing live weight. Instantaneous maintenance requirements increase linearly with increasing live weight when expressed as aspercent of instantaneous feed comunption but expon- entially when expressed in terms of pounds of feed per day. i The graphical solution of the live weight at which the pig-plus-feed cost per pound gain is the least gave a reasonable estimation of the live weight of the animal at which the madman returns, including weaning cost of the animal in terms of pounds of feed, is attained for feed coneueed. ‘ APPLICATION OF THE EQUKPIQV OF THE CURVE W DIHINISHING INCREHENI TO SWINE NUTRITIQJ by Verl E. Headley A THESIS Submitted to the College of Agriculture Michigan State University of Agriculture and Applied Science in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Department of Minal Husbandry 1960 ACKNWIEIDMNTS The author wishes to express his sincere appreciation to Dr. E. R. Miller, under whose guidance, assistance and encouragement this nami- script was prepared. ' ’ Thewriteralsowishestoaclmowledgehisindebudnesstom. J. A. Roofer for his assistance and cooperation in supplying a portion of the naterial necessary for this study. Sincere thanks are extended to Dr. D. E.’ Ullrey for his unfailing interest and encouragenent during the course of this study, and also to hmda Kay Butcher for typing this manuscript. Verl.E. Headley candidate for the degree of Master of Science DISSERTATION: Application of the Equation of the Curve of Diminishing Increment to Swine Nutrition OUTLINE OF STUDIES: major subject: Animal Husbandry (Animal Nutrition) Minor subject: Bio-chemistry' BIOGRAPHICAL ITEMS: Born, January 15, 1927, Eden, Ohio Undergraduate Studies, Tri-State College of Engineering, 1956-1958 Graduate Studies, Michigan State University, 1959-1960 E a Ten Sigma.Eta Honorary Engineering Society TABLE OF CONTENTS INTRODUCTICN....................... REVIEWQ‘LII'ERM‘URE................... History........................ Equationlnprovements................. acquingIntakeIevelswitthuation......... CarparingProteinIavelswitthuation........ ConpaflngBreedswitthuation............ ApplyingEquationtoSteers.............. RecentApplications.................. EXPERDENI‘ALPROCEDURE._................. Derivation of the Equation of Diminishing Increment. . Derivation of Equations for Application. . . . . . . . Solving for Constants A and k by Method of Least sqmaeeeeeeeeeeeeeeeeeeeeeeee Method of Feeding and Mmagenent of Baby Pig on smwetumnmetOOOOOOOOOOOOOOOOOO Composition of Synthetic Milk Diet . . . . . . . . . . Observed Data of Baby Pig on Synthetic Milk Diet . . . Calculations and Adjustments of Constants A and k for Babypignataeeeeeeeeeeeeeeeeeeeee Method of Feeding and Management of Pigs Weaning to Mketeeeeeeeeeeeeeeeeeeeeeeee Coaposition of Typical Ration Used . . . . . . . . . . mutaofPingnWeaningtoMarket. .. .. . Calculations and Adjustment of Constants A and k for Pigsweaning‘toOMBrketeeeeeeeeeeeeeeee Page ‘ONIO‘UIMMH 21 25 26 27 29 36 37 Lo Page Rasmsmmscussxm.................. 1,7 BabyPigCalculations................. 1,7 WeaningtoMarketCalculations............ 56 smxmcmcmsxons.................. 69 BIBLIGRAPHY....................... 71 TABIE or ILLUSTRM‘IONS Page Figure l - General Relationship of Live Weight to Feed Consqution'Conforuingto Curve of Diminishing Increment 114 Figure 2 - weerved Cunulative Efficiency of Feed Utilisation (go) Versus Live Height (w). . . . . . . . 28 Figure 3 - weerved thulative Efficiency of Feed Utilization (go) VON“. u” Weight (')e e e e e e e e 39 Figure I; - Live ht (w) Versus mserved Cumulative Feed Consuaption (fo and Calculated leative Feed muon(tc)seeeeeeeeeeeeeeeeee 50 Figure 5 - Instantaneous Efficiency of Feed Utilisation (Ei)vmuamwwaight(w)eeeeeeeeeeeeee, 51 Figure 6 - Ooserved Cumulative Efficiency of Feed Utilisation (go) and Calculated Cumulative Efficiency of Feed Utilization (13c) Versus Live weight (w). . . . 52 Figure 7 - live weight (w) Versus Cumulative Maintenance Ww< FODZ aflfiunm M CZnfimme4< ruww>wnMM . .flmmfi F>fi>2> bQOOQ HCNu unclubwc—mom 00 40C IOFU >20 3>< 8M moxno€ flow firo0.mn.. Imbcrmws Gem. K.m& ammmmm .000 Dvfirnn>4~oz ow 43m.moc>a~oz on Him chfim om UHZHz~mx~2n ”ZGQMEMZA fio mfiwzm ZCfideHOZ. Zuommm>z mfl>fim CZ~1MJw~d<. 20% 4mm. Dadan LMZZ< X2>CHI\HO vcaucm C rum .mzn arr no _ . . newecm : rum , , . NZU -h1- The num.erical values of C and k are then Obtained by the method of least squares, illustrated in the derivation from equations (21) through (25) and applied as follows: a b 1‘ s bb b1 be '5' ‘fi' 1 - 32e6 Oehol - 31e199 1&076 .1300726 1°17em7h 1 - 6e8 Defi? " hSeh-IB 2190e2h 4.8.1116 2125.328); 1 "' 65e5 0e329 " 615.171 (429er; -21.Sh9h h203e2m5 l - 87. 0.31h - 85.886 7603.8h -27.3808 7&89.2592 l e13h.1 0.2h6 -132.85h 17982.81 -32.9886 17815.721h 1 -157.7 0.236 456.1161. 28869.29 -37.2172 2h67h.3728 1 -180.7 0.220 479.880 32652.1.9 -39.7Sho 32h32.0360 1 -202.8 0.211 -201.589 81127.5h ~h2.7908 39368.2h92 - e e " e2 3 e "' 0 3 0 3 or 01‘ 01‘ or 01‘ 02‘ 01‘ (“)1 (ab)1 (81h (”)1 (bb)1 (bl)1 (bs)1 Check: (aa)1 + (ab)1 * (al)1 ' (as)1 9 - 1017.9 +' 2.627 - ' 1006.27 (ml + (bb)1 + (bl); - (ten -1017.9 + 110989.77 - 268.1366 - 11727073331. Calculations for the first estimate of k, c, and 11. k-ébl)“ 016w}; k+§algls 1-0/1: 2 ‘3 l as 1 (bb) - (0b) - ah ab 3 (b1) - (b1) - (ab)1 (a1) 2 1 1.3.3:); 2 1 7.2534 1The equationsused here-for calculating k and C are identical to the previous dgrived equations (2b) and (25), for example (b1)1 is m to “Es ' ~172- (bb)2 - 110,989.77 - 11,911,221 - + 28,865.28 9 (bl-)2 '-26h.1366 - 5-1017e23 S2e6272 . + 32e9771 ga,§§;e58 C -{.1017.9)_(0.0011h211h8§) + 2.627 - 0.1:2109982 9 9 A " 0. 1 82 " 3&.S9h092 $132388 ' The first estimates of A and l: have now been calculated. The next step is to adjust the values of these constants A and k in such a way as to reduce to a minimum the sum of the squares of the differences between the observed cumulative efficiency and the calculated cumulative efficiency. This is done exactly as shown on the baby pig data previously, that is through the application of the method of least squares Qplied to equation (26) by the following procedure: Calculations for fc for wi - 26.6, A - 368.59h092, k - 0.0011112171186, (m1) - 31.1.998092 w w (m) ((1331 108%. £°-1081(A—§: ' 9 k m 26.6 3141391092 1.0000000 0.000000000 ‘ 0.000000 38.6 329.99h092 1.0363613 0.015513006 31.266190 55.0 313.591.092 1.0905629 0.0370091 75.881966 75.9 292.69h092 1.1688352 0.067601021. ' 136.21.8673 98.5 270.09h092 1.2662035 0.102501225 2065891112 122. 21.6.1911092 1.3891239 ~ 0.1h2738h09 7 287.6867748 11.5.8 222-.7911092 1.5350232 04861161796 3754111518 169.6 198.9914092 1.7186113 0.235179718 h7h.000577 191.8 176. 798092 1.930201: 0.28655sz8 577.51.11.90 213.7 15113917092 2.2079221 0.31.39811120 693.291.537 Calculations for go, go: and (5° -.§c) as follows: -1.3- ' I. to H1 Ea.“ c. 0-(EO-Ec) c -551' g 27:1 c c 26.6 00.0 00.000000 00.0 M 38.6 29.9 31.266190 12.0 0.1.013378 0.3838012 0.0175366 55.0 72.2 75.881966 28.1. 0.3933518 0.3782655 0.0190863 75.9 135-8 136.21.8673 119.3 0.3630339 0.3618 3814 0.0011955 98.5 207.7 206.5891112 71.9 0.31.61721. 0.31.80333 0.0018609 122.1. 292.2 287.6867h8 95.8 0.3278576 0.3330011 0.0051135 11.5.8 387.1. 375.111.518 119.2 0.3076923 0.3177670 0.0100717 169.6 1.88.1 h71..000577 11.3.0 0.2929728“ 0.3016871. 0.0087116 191.8 588.9 577.51.11.90 165.2 0.2805230 0.28601400 0.0055170 213.7 693.8 693.29h537 187.1 0.2696713 0.2698709‘ 0.000196h Calculations for gc/k and (Ec)2/k(lfifli)(lev) 8.x]. (8.)2 we.) (1...) (892 ° ° WHYUHJ 335.91.61.30 0.111730336 128.932216 0.001111211868 327599809 0.11.007h66 122.521.561 .0.00111.32371. 316.721809 0.13092703 111..358707 0.00111.1.8803 301.301.9111 0.12112718 105.52861.6 0.0011178133 291.1.801119 0.11088973 96.190661 0.0011528118 278.11.55115 0.1009758? 87.01.8031 0.0011600017 261..070861. 0.09101529 77.71.9117 0.0011706279 250.371.1193 . 0.08181888 69.075310 0.001181.1.B75 236.2211172 0.07283033 60.518776 0.001203h336 eeowpsdom .23 us find ma page: no: one hog .eeaseee Henna one seems. ease: sees: sheen. meanness an esflee no es. ash 63»an 0.33 Agdoolmv usages moons on» sand» mane dHH n emaaameooooo.o+ moaemauaoooo.o amomawa.n+ aeuommao.n «aaaac.aoosea ammonuoooooo.ou ommwmeaaooooo.o oaaneao.o. emeawam~.o mmcmmm.oommm semaooo.ou enmamomaoo.o ~a4-~.om~. mawammoooooo.o. aoaomoeaooooo.o Heamamn.a. pamemeem.o msaewm.ame~6 cadmmoo.o. macaeoaaoo.o neaeam.om~ ammeomoaoooo.o. meemo~maooooo.o o-~aon.~- mammaaom.o 4H~H~4.mmaae eaaamoo.o. mameoaaaoo.o accoao.ae~ aseaaaaooooo.o. amoememaooooo.o mummwom.~- onmaemwm.o momqaa.aemaa aeaooao.o- aaooooaaoo.o mammaa.ma~ megawamooooo.on omamommaooooo.o Hemmmea.au chamoemm.o Howaae.oomam mnaamoo.o. magnumaaoo.o assom:.em~ oommmamooooo.on mmaeaamaooooo.o eaamaom.on camaaman.o awmeoo.aommm moccaoo.on mnemaeaaoo.o asunoa.aon 42838686. 82328686 8.826. $86806 832.38H mmad86. 8844286 «8.856% «anemmamoooo.o+ aaaaeomaooooo.o ooeo~m~.6+ ammwmaam.o aaamam.fl~maon neaoaao.o+ samwnaaaoo.o eoemem.-m ammmmoomoooo.o+ meawmomaooooo.o mmmmamm.m+ noaammon.o aemmom.ammwaa oenmaao.o+ momawaaeoo.o omaeaa.mnn n31<~ws=1<~x 1m: o «1.5 em .3 ab .7 as ea H p e “omega.“ ee Hacksaw sewed no e058. on... he eopegoaee on see m ens Us we need; 83.35.: 05. 605.396 so: needs» Scenes: 05 and» 4:5- Gslculations for the correction values 0C and Q 9 '(b1)23 0C- 4.»); e + (a); “1 . ( (uh); I 7 “51 (bb)2 - (bb)1 - gab}; (a); 3 (01)2 - (01h - (ebh (:1); as 1 an 1 ‘ (bb)2 - 0.000012136705 .. S250l25629222 - 0.000000202591 7 ,007. 991 2 (01);, - 0.000006511726 - g3,0%%E%;2 éfizggoan ’ e --0.000009000162h 8 - - gwm?!‘ - - hu.u25282 ~ 0C ' W ” W ,00 . 991 2 , . 172 . 0.00018071766 I Since 8 is the correction for A. and etc is the correction for k, the first adjusted values for A end 1: then calculated as follows: 368591092 0.001lb2hh86 Adjusted A 32 .1 1 Adjusted k - . 13231 3 Using these adjusted values of A and 1:, new velues were calculated for :6, 13c, and (go - EC) end the same procedure was used for making the second adjustment (en adJustnent of the previously adjusted values of A and 1:). These adjustments were continued until the correction values of e endoC were small in comparison to A and 1:. Three adjustments were required for A and k on the data pertaining to the pig from weaning to nerket. Adjustment results were as follows: First estimate After first adjustnent After second adjustment After third adjustment Spillnsn ratio, 4:6- ...é... 368 . 5911092 326.168810 331.814th0 331.678085 e'k - 0.99868273 k 0.0011h2hh86h 0.00132316630 0.00131h95825 0.0013151377h .-h7- RESULTS AND DISCUSSIGJ Babz Pig Calculations _ In reference to Table I the calculated cumulative feed comuaption (fc) of the baby pig on a synthetic nilk diet conforms with fair accuracy to the observed values (see Figure 1;). The instantaneous efficiency of feed utilization (E1) decreases linearly with increasing live weight (see Figure 5). The, instantaneous feed conversion (F1) increases with increasing live weight. Cumlative efficiency of feed utilisation decreases linearly with increasing live weight (see Figure 6) and the reciprocal of (gc) or cumulative efficiency of feed conversion (13¢) as would be expected showed more feed consumed per pound gain as live weight increased. In reference to Table II the cumulative maintenance requirement (Me) was found to increase exponentially with increasing live weight (see Figures 7 and 8). The maintenance requirement between any two live weights was shown to have an increasing effect as live weight of the baby pig increased. Also the instantaneous maintenance requirement (M1) expressed as a percent of instantaneous feed consumption was found to increase linearly with increasing live weight (see Figure 9). Note: As previously illustrated, maintenance requirements as applied in this study and as defined by the equation include all feed consumed exclusive of that which is utilized for protein storage (growth). -h8- l‘lll $53386 .. a Seamuem .. 3 ‘l moa-a~n.a o~m4m4m~.o «Hammmom.a memoumom.o caumpw.m~ oo.m~ $283.4 Sfiofle6 endgame; oflmfioed 383.3 8.8 $334 Seaweed «canine; sanded 423.2 8.: Sign; . 9329.86 omemmnemé $533.6 8286 8:: 5.39884 3.8436 enigma; 803386 8886 84a ommacpsm.o mm:~mm~o.a bmmmmauo.a mmaaamvm.o mmoaom.~ oo.o amnoemme6 «88864 888.8 aim LEI on on Ari: 3...; 3-4: I 0.0% I 0m I HM I H.” ’ encapesuo ee>wnen noun noanne>cco seem new honoaoauue obaaeassso ene.nsoonwanuneeH .aa scum neaanon uncapdsuo henna new Hanoaeuona madnnacasao on» we convenes one no naaneem 83 ea: one-sedan .- do and .33 23 e0 33 3 83.5 30323 we 398m H Danna hnamundmod a a 3888.8 .. «a 888.8 888.8 4888.“ 288:8. 883.8 8.88 4.88.5 888.8 33988 84856 388.8 8.8 888.8 888.3 28084 8888...” 3838.2 . 8.3 888.8 888.8 888.24 28.8.... mS8o.a 8;: 888.8 388.8 «882.6 «.3884 8886 85 0., 538.3 888 .8 38846 38346 , 8888 86 .w Isis... 5886.2 68886 88866 688.8 mum 8a a d .. N , . nlllb. . LP 3. w 388.39 was”. 8” e .. 8%an .. :1: 3... ea E IN mad I H 3 Gene «0 a new WOO 800m no a 9: «z .. man-Hem .. o: incoapesvo enhance scnu 3558.300." cognac: 333350 was 3853.33 83 can eeeefihnm . do 5 Pan 33 no 33 3 sundae 3333 we 33 HH 0.3:. -50. 25 " 20 w (leo) 15 10 S o 5 ' 10 15 20 25 f0 and {C (3)8.) Figure 1;. Live Weight (1:) Versus Observed Cumulative'Fecd Consumption (f0) and Calculated Cumulative Feed Conannption (to). , -51. 1.2 f 1.0 F 0.8 ‘ 0.1: #- 0.2 F l l 0 5L i0 15 20 W (use) Figure 5. Instantaneous Efficiency of Feed Utilization (E1) Versus Live Weight (w). .52.. 1.2 [ 1.0 ' dE 13°“cc 0.6 ‘" ooh F' 0.2 F i I 4 AL 0 5 10 15 20 w (1ba.) Figuro 6. Oosemd melativn Efficimcy of Food Utilization (Eo)lnd Calculated Cumulative Efficiancy of Food nunzauon°(sc) Versus Live Weight (1:) . -53- 25 - 20 - W (1138.) 15 — 10 L. S o 2 h 6 5 173 4 “c (1138.) Figure 7. Live Weight (w) Versus Cumulative Maintenance Requirement (Me) . -51.. 25 F N C re 20 - V (1138.) 15 - 10 ’ 5 0 S 10 15 20 25 to and MC (1be.) Figure 8. Live Weight (17) Versus Cumulative Feed Consumption (re) and Cumulative Maintenance Raduiremnt (Me) . -55- ho e Mi (as X 01 feed) 30- 20... l l L l J o 5 ‘ 10 15 20 25 w (lbae) Figure 9. Instantaneoue Maintenance Requirement (M1) as Percent of Inetantaneeua Feed Consumption Verena Live Weight (3). -56. Nagging to Market Calculations In reference to Table III the calculated cunnflative feed consuqation (fc) of swine from weaning to market conforms with a high degree of accuracy to the observed values (see Figure 10). likewise, the calculated cumula- tive feed commtion (fc) conforms with accm'acy to National Research Council (N.R.C.) standards (Beeson £93, 1959) as shown in Figure ll. The instantaneous efficiency of feed utilisation (31) decreases linearly with increasing live weight and conpares to N.R.C. standards to approxi- mately 160 pounds live weight. Figure 12 indicates that swine from 160 to 200 pounds live weight may not be as efficient as N.R.C. standards designate. The instantaneous feed conversion (F1) increases with in- creasing live weight. Cumulative efficiency of feed utilization decreases linearly with increasing live weight (see Figure 13) and the reciprocal of (13c) or cumulative efficiency of feed conversion (13c) showed more feed consumed per pound gain. as live weight increased. In reference to Table IV the cumulative maintenance requirement (Me) was found to increase exponentially with increasing live weight (see Figures. 11; and 15). The maintenance requirement between any two live weights was shown to increase as the live weight of the animal increased. Also instantaneous maintenance requirement (Mi) expressed as a percent of the instantaneous feed consmption was found to increase linearly with increasing live weight (see Figure 16). In reference to Table v the calculated daily feed consunption conpares very well with N.R.c. standards. The instantaneous daily main- tenance requirements indicate that the pounds of feed required for daily maintenance increases with increasing live weight. It should be pointed -57- out that the instantaneous daily food consumtion and maintenance requirements were obtained by the tangential method from the curves of cumulative feed consumption(fc) versus days of age and cumulative main- tenance requirements versus days of . age at points corresponding to spec- ified live weights. The pounds of feed available for growth were obtained by the diff- erence between daily feed consunption and daily maintenance requirement. It should also be pointed out that calculations indicate the ratio between daily gain and pounds of feed per day available for growth bears a reason- ably constant relationship. Plotting as in Figure 17 yields the graphical solution of the live weight at which pig-plus-feed costs per pound live weight gain are the least. This gave a reasonable estimation of the live weight of the animal at which madame returns, including weaning cost of the animal in terms of pounds of feed, were attained for feed consumed. -53. 5.888686 .. s . member—Hm a 1n 88.386 28.3.6.6 8.3.5.6 85.856 3486.80 . 6.86. 38336 4.35686 8.88%.: 88886 masheom 6.3.6 88836 88886 636886 88686.6 888.48 0.86 6.84386 088686 28836 383.66 9886.86 8mm." 68836 6838m6 8888.8 88386 48:88... o.§ «$68.96 683686 638686 488886 868.8." o5. 83886 33886 863636 _ 984.886 «868.8 0.8 . 836.36 88836 o88o.8 ed... 35 u a ow I .. < e. a .o IsL j rJLu e. - a. E I O“ I 0m I .Hh I flu I 0H ' .afiagve enhance Ion.“ defiance—hoe been one hounded: spanned—.50 use magma” ceases 8. 655.: none omen no 3.8 3 cognac accede—am no coined . HHH 0683a . 3 done coarse escapees. noneo one sandstone geodesic one we convenes one. no 398m 5.2.5285 .. x 883.3 838.8 388.2. _ amamé... . 3.38.5 0.86 888.3 6862.6.“ 386.8 3.6363 mfiaohéom o5." $88.3 9.38.3 «6.8.9.3 2.5:...3 8825mm o6? 888.2 4.58.: 423.8 8382. 8886.86 0.66 933.8 3.36.8 88.3.8 288.2 32.3.86 0.86 48893 866.366 8.38.3 8.38.8 86862 0.2. 38% :3 538.3 538;. $2.84. 358.8 0.8 . .6836 0880.8 888.8 08898 0.8 an A a m Howm~ SA n F I