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Daphnia (Prey)
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Ea = 0.04 (0.02)Temperature (¡C)
ln (attack rate)
(c)101520253035!2!101!!!!!!Ea = 0.67 (0.12) eVEa = 0.58 (0.06) eV!0.20.00.10.20.3ln (relative growth rate)
(d)Temperature (¡C)
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Prey density
Temperature                   
Active
Sit!and!wait
Temperature                   
Active
Sit!and!wait
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T=;@Jr?=K@JFGF@<9C
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!!!!!!!!!!!!!"#!X__!!B(%7'/!8,-!4'&;&')(&2!&E!%/2/'1!(2!/1:*!A1)/'0*/3!C/$&2%(2%!)&!/1:*!)'1()!
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"@C=@KG=;9S!'B=J=!G9V9!B9I=!K;FE9;FC8!JE9CC!D@?8!JFW=JO!EFV=?!;B=@KBFC8O!K;FE9;FC8!A@@C
T>9;E!
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T![R!}"Z!K;=L=;=<A=JO!9<?!DF
TE:CGFI@CGF<=!CFL=
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GB9G!B9I=!K;FE9;FC8!E=?F:E!G@!C9;H=!D@?8!JFW=J
O!?=K@JFGF@<9C!;B=@KBFC8O!A@@C
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8GB=;E9C!
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T![R!}"Z!K;=L=;=<A=O!9<?!:<FI@CGF<=!9<?!J=EFI@CGF<=!CFL=
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!!!!!!!!!!!!!!!!!!!!X_^!!B(%7'/!88-!M&/EE(:(/2)!/0)(+1)/!F+/3(12!P![?c!:'/3(C$/!(2)/'@1$0G!E&'!/1:*!;'/3(:)&'!
@1'(1C$/!(2!0;1)(1$!+&3/$0!A()*!
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!+;9<H=!K@F<GJ!F<?FA9G=!
A@=LLFAF=<GJ!>FGB!^]i!A;=?FDC=!F<G=;I9CJ!<@G!@I=;C9KKF<H!W=;@S!
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!0.000.250.500.751.00Baseflow
Cultivated
Forested
SlopeSpaceTemp.
Precip.
Coefficient estimate!Genus richness
EPT richness
Functional richness
Rao's Q!X`R!!B(%7'/!8<-!M&''/$1)(&2!:&/EE(:(/2)0!E&'!;'/3(:)&'!@1'(1C$/0!E'&+!E&7')*!:&'2/'!121$I0(0!A()*!
)'1()!;'&E($/!%'&7;0-!
+;9<H=!K@F<GJ!F<?FA9G=!A@=LLFAF=<GJ!GB9G!>=;=!JFH<FLFA9<G!9LG=;!9?P:JGE=<G!@L!
KTI9C:=JS!
3:;KC=!K@F<GJ!F<?FA9G=!A@=LLFAF=<GJ!GB9G!9;=!<@G!JFH<
FLFA9<GS!
(;;@;!D9;J!9;=!@EFGG=?!L@;!
AC9;FG8S!
!!!!!!!!!!!!!!!!!!!!!!!!!!0.02!0.010.000.010.02Baseflow
Cultivated
Forested
SlopeTemp.
Precip.
Coefficient estimate!Trait profile 1
Trait profile 2
Trait profile 3
!X`X!!B(%7'/!8=-!M&''/$1)(&2!:&/EE(:(/2)0!E&'!;'/3(:)&'!@1'(1C$/0!E'&+!E&7')*!:&'2/'!121$I0(0!A()*!
(23(@(371$!)'1()0-
!);@E!G@K!G@!D@GG@E2!;B=@KBFC8O!GB=;E9C!K;=L=;=<A=O!9<?!I@CGF<FJE!Y<:ED=;!@L!
H=<=;9GF@<J!K=;!8=9;ZS!4F
TE:CGFI@CGF<=!FJ!G>@!@;!E@;=!H=<=;9GF@<J!K=;!8=9;O!:<FI@CGF<=!FJ!@<=!
!!!!!!!0.10.00.1Baseflow
Cultivated
Forested
SlopeTemp.
Precip.
Coefficient estimate!DepositionalDepositional!erosionalErosional!!!!!!!0.2!0.10.00.10.2Baseflow
Cultivated
Forested
SlopeTemp.
Precip.
Coefficient estimate!< 5¼C0!15¼C5!30¼C15!30¼C>30¼C!!!!!!!0.08!0.040.000.04Baseflow
Cultivated
Forested
SlopeTemp.
Precip.
Coefficient estimate!Bi!multivoltine
Univoltine
Semivoltine
!X`Q!B(%7'/!8=-!
F:&2)N3G
!H=<=;9GF@<!K=;!8=9;O!9<?!J=EFI@CGF<=!FJ!L=>=;!GB9<!@<=!H=<=;9GF@<!K=;!8=9;S!
(;;@;!D9;J!9;=!@EFGG=?!L@;!AC9;FG8S!
!!!!!!!!!!!!!!!!!!!!!!X`[!!B(%7'/!8?-!O/12!:$(+1)/!0/20()(@()I!0:&'/!E&'!(20/:)!3(@/'0()I!+/107'/0!F)&;G!123!)'1()0!123!
)'1()!;'&E($/!%'&7;0-
!';9FG!9DD;=IF9GF@<J!9;=2!;B=@KBFC8O!5=K@r?=K@JFGF@<9CO!5=K@
T=;@Jr?=K@JFGF@<9C
T=;@JF@<9CO!(;@Jr=;@JF@<9Cn!6@CGF<FJEO!4F
TE:CGFrDF
TE:CGFI@CGF<=!
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K=;!8=9;ZO!1<Fr:<FI@CGF<=!YX!H=<=;9GF@<!K=;!8=9;ZO!*=EFrJ=EFI@CGF<=!YuX!H=<=;9GF@<!K=;!8=9;ZS
!!!!!!!!!!!!!!!X`\!!!B(%7'/!8V-!4'&;&')(&2!&E!%/2/'1!(2!/1:*!A1)/'0*/3!:$100(E(/3!10!*(%*$I!0/20()(@()I!)&!:$(+1)/
-!"CFE9G=!J=<JFGFIFG8!FJ!A9CA:C9G=?!9AA@;?F<H!G@!J=<JFGFIFG8!@L!
=9AB!H=<:Jg!
;B=@KBFC8O!GB=;E9C!
K;=L=;=<A=O!9<?!I@CGF<FJE!G;9FGJ!YC=LGZ!@;!G;9FG!K;@LFC=!H;@:KJ!Y;FHBGZS
!!!!!!!!!!!!X`]!!!!!!!!!!!!!4$4#$+0-&3.7
!!!!!!!!!!!!X`f!4$4#$+0-&3.7
!!&CC9<O!dS5S!QRR\S!
#9<?JA9K=J!9<?!;FI=;JA9K=J2!GB=!F<LC:=<A=!@L!C9<?!:J=!@<!JG;=9E!=A@J8JG=EJS!
&<<:9C!;=IF=>!@L!=A@C@H8O!=I@C:GF@<O!9<?!J8JG=E9GFAJ
![]2Q]_
NQ`\S!!&JKF<O!'S/S.SO!aS!aB9EFJO!'SdS!%9GGB=>JO!&S%S!%FC<=;O!%SdS!+g"9CC9HB9<O!%S!';FEE=;O!0S!
/@@?>9;?O!9<?!%S(S!#=?H=;S!
QRX^S!(VG;=E=!?;@:HBG!K:JB=J!JG;=9E!F<I=;G=D;9G=!A@EE:<FGF=J!
@I=;!L:<AGF@<9C!GB;=JB@C?JS!0C@D9C!"B9<H=!4F@C@H8!Q]2Q[R
NQ\\S!!49CF9<O!(S6SO!.S!*=H=;JO!"S!#„I’M:=O!9<?!aS!%9;G=<JS!QRR`S!'B=!);=JB>9G=;!&<FE9C!5FI=;JFG8!
&JJ=JJE=<G2!9<!@I=;IF=>!@L!GB=!;=J:CGJS!.
8?;@DF@C@HF9!]^]2fQ_
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QRRRS!%=9J:;F<H!GB=!9GG9F<E=<G!@L!DF@C@HFA9C!F<G=H;FG8!F<!GB=!1*&2!9!A;FGFA9C!=C=E=<G!@L!
=A@C@HFA9C!F<G=H;FG8S!39H=J!\][
N\f\!*+!%S!d:<H>F;
GBO!*S!%:B9;O!9<?!*S!*ABE:GWO!=?FG@;JS!
&JJ=JJF<H!GB=!(A@C@HFA9C!$<G=H;FG8!@L!-:<<F<H!/9G=;JS!*K;F<H=;!,=GB=;C9<?JO!5@;?;=ABGS
!!49;<=GGO!'S3SO!5S/S!3F=;A=O!.S0S!.F?9CH@O!"S!4@<LFCJO!4S5S!*9<G=;O!'S!59JO!0S!49C9O!&S/S!/@@?
O!=G!9C
S!QRR`S!.:E9<
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!F<!GB=!.8?;@C@H8!@L!GB=!/=JG=;<!1<FG=?!*G9G=JS!*AF=<A=!
[X^2XR`R
NXR`[S!!49VG=;O!"S6SO!aS5S!)9:JABO!9<?!/S!"9;C!*9:<?=;JS!QRR]S!'9<HC=?!>=DJ2!;=AFK;@A9C!LC@>J!@L!
F<I=;G=D;9G=!K;=8!CF<U!JG;=9EJ!9<?!;FK9;F9<!W@<=J2!3;=8!J:DJF?F=J!CF<U!JG;=9E!9<?!
;FK9;F9<!L@@?!
>=DJS!);=JB>9G=;!4F@C@H8!]R2QRX
NQQRS!!4@<9?9O!,SO!*S!5@C„?=AO!9<?!4S!*G9GW<=;S!QRR_S!'9V@<@EFA!9<?!DF@C@HFA9C!G;9FG!?FLL=;=<A=J!@L!
JG;=9E!E9A;@F<I=;G=D;9G=!A@EE:<FGF=J!D=G>==<!E=?FG=;;9<=9<!9<?!G=EK=;9G=!;=HF@<J2!
FEKCFA9GF@<J!L@;!L:G:;=!ACFE9
GFA!JA=<9;F@JS!0C@D9C!"B9<H=!4F@C@H8!X[2Xf]`
NXf_XS!!4@<9?9O!,SO!,S!3;9GO!6S.S!-=JBO!9<?!4S!*G9GW<=;S!QRRfS!
5=I=C@KE=<GJ!F<!9M:9GFA!F<J=AG!
DF@E@<FG@;F<H
2!&!"@EK9;9GFI=!&<9C8JFJ!@L!-=A=<G!&KK;@9AB=JS!&<<:9C!-=IF=>!@L!(<G@E@C@H8!
]X2\^]
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0FCC=GO!9<?!3S!#=H=<?;=S!QRXXS!,:E=;FA9C!(A@C@H8!>FGB!-S!*K;F<H=;
T6=;C9HO!,=>!
7@;US
!!4@GG9
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E:CGFKC=!G;9FGJS!d@:;<9C!@L!I=H=G9GF@<!JAF=<A=!Xf2][[
N]\RS!!4;9H9O!dSO!"SdS)S!4
;99UO!/S!'B:FCC=;O!9<?!*S!5;98S!QRX`S!$<G=H;9GF<H!JK9GF9C!9<?!KB8C@H=<=GFA!
F<L@;E9GF@<!F<!GB=!L@:;GB{A@;<=;!9<9C8JFJ!G@!G=JG!G;9FG
N=<IF;@<E=<G!;=C9GF@<JBFKJS!(A@C@H8!
^^2Qff_
NQf_\S!!!X`_!4;9:<=O!(SO!+S!-FABG=;O!5S!*y<?H=;9GBO!9<?!)S!*:BCF<HS!QRR`S!6@CGF<FJE!LC=V
FDFCFG8!@L!9!;FI=;F<=!
?;9H@<LC8!9C@<H!GB=;E9C!H;9?F=<GJS!0C@D9C!"B9<H=!4F@C@H8!X\2\_R
N\`QS!"B9@O!&SO!9<?!#S!d@JGS!QRXQS!"@I=;9H=
TD9J=?!;9;=L9AGF@<!9<?!=VG;9K@C9GF@<2!JG9<?9;?FWF<H!
J9EKC=J!D8!A@EKC=G=<=JJ!;9GB=;!GB9<!JFW=S!(A@C@H8!^[2Q][[
NQ]\_S!!"B=JJE9<O!4
S"S!QRXQS!4F@C@HFA9C!G;9FGJ!K;=?FAG!JBFLGJ!F<!H=@H;9KBFA9C!;9<H=J!@L!L;=JB>9G=;!
F<I=;G=D;9G=J!?:;F<H!ACFE9GFA!>9;EF<H!9<?!?;8F<H2!$<I=;G=D;9G=!G;9FGJ!9<?!ACFE9G=
T;=C9G=?!;9<H=!
JBFLGJS!d@:;<9C!@L!4F@H=@H;9KB8![^2^]_
N^f^S!!"@<GFO!#SO!&S!*ABEF?G
TaC@FD=;O!0S!0
;=<@:FCC=GO!9<?!/S!0;9LS!QRX\S!&!G;9FG
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