I"? I II I I I II II I I I I I I II . II‘ I.I., I I II I l STUDIES ON CURLY COAT, GROWTH, MULTIPLE BIRTH AND COLOR OF YOUNG HORSES Thesis for "the Degree of M. S. MICHIGAN STATE COLLEGE Leonard H. BIakesIee I 941 . , o p: ‘putu Ir . 1.140 {3:11.0110I-11w; I nt..rW-Inc.o 4 . ..I. ..I .. . t ' . . . u I O! a . DI . alvo. . w. . I I AI... H. .c. IflmownlIn.Il.. A . p v . n . I ..a ....I. 10v.” v.' I I I . I . .11 In . . .. . Jyntr IA .. 0. g .vIl . IHID . . . . . .1. .0, I . . I I . a .|,I . ., I. 4“ .. Ir .Ju1._ ...._.dafia.drt:u1rf r: o.nr .4. . _.III W' ‘ “it. \ ’1“ «1’»; ~ ' 1|..- h'),1.:" '4 I {W w 0 ' . '. ‘ f I I' l 3"“ Liv PLACE IN RETURN BOX to remove this checkout from your record. TO AVOID FINES return on or before date due. DATE DUE DATE DUE DATE DUE ‘—'MRGIL. c oewfi 1/98 C‘JCIRC/DateDue.p65-p.14 .. _' 3% 1. AI.” . .4 “I ‘2 : [b 1,} ‘1' - tr»; ’P'Ari, . In :1. (II. ,I STUDIES ON CURLY COAT , GROWTH, MULTIPLE BIRTH AND 001.08 OF YOUNG HORSES Thesis for Degree 0!. Heater of Science Michigan State College Leonard H. fiakeelee 19M u.’\1umfl‘ri .Iluer.-JLlu«. , THESIi . II STUDIES ON CURLI COAT, GROWTH, MULTIPLE BIRTH AND COLOR OF IOUNG HORSES A THESIS SUBMITTED TO THE GRADUATE SCHOOL OF TECHIGAN STATE COLLEGE OF AGRICULTURE AND APPLIED SCIENCE BI LEONARD H. BLAKESLEE IN PARTIAL PULPILIAIENT OF THE REQUIREMENTS FOR THE DEGREE OP MASTER OF SCIENCE IN ZOOLOGT MAJOR SUBJECT: GENETICS EAST LANSING 19141 Acknowledgment Sincere appreciation is hereby expressed to Dr. H. R. Hunt, Head of the Department of Zoology for council and criticisms, especially'with the genetic phase of this manuscript, and to Dr. W; D. Baten, Associate Professor of’lathematics, for generous assistance in the mathematical analysis. The sincere appreciation of the author is also extended to Professor G. A. Brown, Head of the Animal Hquand y Department, and Professor R. 8. Hudson, Superintendent of the Farm and Horse Depart- ment, for time and materials which made this study possible. 142797 Tgplj of Contents INTRODUCTION............................................... 1 PART I - Curly Coat of Horses............................. 3 Review of Literature............................. 3 Occurrence of Curly Coat in Horses............... 3 Curly and Smooth-C oated Foals of Similar BreedmgOOOCOCOOOOOI00....OOOCOOCOOOCOOOOCOCCOO 9 PART II - Growth Studies in Foals..........................1O Introduction and Review of Literature............10 Problem and Method of Procedure..................13 Presentation of Data.............................l7 Growth Rate Comparisons of Breeds and Sexes......29 PART III - Multiple Births in Horses.......................M Data on the College Purebred Herd................h5 PART IV -@lor DevelOpment in Draft Foals..................lo8 Review of Literature................Human...“ Color of Foals at Birth..........................l+9 SIMARY....................................................51 BBLIWHIOOOOOOOOOO000......0.000000000000000...00.00.0052 APPWIXOOCOOOOOOOOOOOO...0.00.00...0.0.000...00.000.00.00055 .1 d r VA}. LIN.“ .bIrLGJI 5.5». I hillQ; amazes 1. 2. 3. h. 5. 6. 7. 8. 9. 10. 12. 13. 15. 16. 17. 18. 19. Page Colonel Last - A Curly Colt as a Foal............. ‘6 Colonel Laet as a Yearling........................ 7 Metric Measuring Instruments...................... 12 Measurements Recorded............................. 1A Graph of Average Weights and Body Depths. . . . . . . . . . l9 Graph of Average Weights and Height at Withers . . . . 20 Growth Curve for Averages of Body Depths. . . . . . . . . . 21 Growth Curve for Averages of Height at Withers. ’. . . 22 Graph of Average Weights and Body Lengths..... .... 23 Growth Curve for Averages of Body Lengths. . . . . . . . . 21; Graph of Averages “of Weights, Heart Girth and PrediCted WeigltSOeooeeoeoeeeooeeeeeoeeooeeeoee 25 Growth Curve Famed by Averages of Heart Girths... 26 Growth Gn've Formed by Averages of Weights.... ..... 27 Hip ldth Growth Curve of Belgians and Percherons.. 36 Bodv Depth Growth Curve of Belgians and Percherons 37 Heart Girth Growth Curve of Belgians and PercheronsBB Shoulder Width Growth Curve of Belgians and PerCheronBOOOCOOOOOOOOOOOOOOOOOOOOOOOOOOOOOO... 39 Height of Withers Growth Curve of Belgians and PerCheron8...0...’..........0......0......QQOQQ ho Chest Floor to Ground Growth Curve of Belgians and PerCheronsoeoeoeeeeeeeeeeee0000000000000... 1‘]- Body Length Growth Curve of Belgians and Percheronsaz Weight Growth Curve of Belgians and Percherons.... A3 calet mdmnEll-yFoal-80.00.00.000OOOOOOOOOOOO... 1’6 ix}. V .. 137$ bl-.. «5.91 Tame Eggs 1. Curly and Smooth Foals of Similar Breeding......... 9 2. Average Weights and Measurements of AB Foals....... l6 3. Average Increases in Body Measurements and Weights. 18 A. Estimated Weights of Draft Horses from Heart Girth neasmentBOOOOOOOOOOOOOOOOOCOOOOOOOOOOOOO m 5. Average Measurements and Weights of Belgian and PerCheron “ItSOOCOOOOOOOOOOOOOOOOOOOOOOOCO...O... 3h 6. Average Measurements and Weights of Belgian and PerCherOn Fillies.eeooeeeeoooeeeeoeeeoooooooeoooo 35 7. Foaling Sumary at Michigan State C allege. . . . . . . . . . A7 8. Foaling Record l936 of Belgians and Percherons. . . . . 56 9. Foaling Record 1937 of Percherons. . . . . . . . . . . . . . . . . . 57 10. Foaling Record 1937 Belgians....................... 58 ll. Foaling Record 1938 of Belgians and Percherons. . . . . 59 12. Foaling Record 1938 of Belgians and Percherons..... 60 Pediggees 1. A le Calt Foaled in 19310000000000.000000900coco 61 2. Colonel Laet - A Curly Colt Foaled in 1932......... 62 3. Queenet - A Curly Filly Foaled in 1932............. 63 A. A Curly Colt Foaled in 1933 6a 5. A (Curly Colt Foaled in 1932 65 6. A Curly Colt Foaled in 1939........................ 66 7. A Curly Colt Foaled in 1939........................ 67 8. A Curly Colt Foaled in 19h0........................ 68 INTRODUCTION .Iany noteworthy contributions have been made to the genetic field since Handel's law of inheritance was made known to the world. lendel's work, and that of other eminent genetists, served not only as an incentive to workers, but also as a tool in animal and plant breeding. While much progress has been made, there still remains a vast and unexplored field for study in animal genetics and all of its related subjects. The obstacles encountered in livestock genetic investigations make progress difficult. A few of these obstacles are slow rate of increase, influence of environment, control of environment, lack of uniform.methods of evaluat- ing offspring, need for uniform records, and the economic standards in- volved. The outstanding obstacle in studying the genetics of the horse is its slow rate of increase which greatly delays the accumulation of the necessary information. Furthermore, horses are not considered.mature until they are four years old, in spite of the fact that many reach mature weight earlier than this age. Fully cognizant of these facts, the writer felt when this study was initiated that an Opportunity presented itself which could not be overlooked. The horse herd owned by Michigan State College is unique in its origin and deve10pment, making it a desirable basis for the studies reported in the following pages. This herd, composed of Belgian and Percheron draft horses, was deve10ped largely from.the offspring of two outstanding females. One, a Belgian called.Pervenche 9030, is twenty years old. This mare, her daughter, and their offspring, make up most of the herd of Belgian mares maintained at.lichigan State College. Pervenche was selected as a foal because of her very desirable type. After her purchase in 1922, she made an -2- outstanding show record which, supported by her record as a brood mare, justified her selection as a foundation animal. Nine of her female off- spring or their descendants are now in the College herd. These nine mares are the get of Range Line Phoenix 12808, Preston Wolfe 11.607, Success de Cognebeau 17015, and Ginger 18711. The other mare, Coreen, a grey Percheron, was equally as famous. This mare was foaled in 1911. and was purchased by the College in 1923. Since coming into the College herd, she has completed an outstanding record, producing twenty foals during her twenty-five years. Of this number, sixteen were strong, living foals. One pair of twins and one single were aborted. The twentieth foal was dropped during her last year, but did not live. The present Percheron herd is com- posed largely of Coreen's female offspring or their descendants, twelve in nunber. These mares are the get of such sires as Treviso 151.392., Sir Laet 190277, Oak Forest Echo 18351.0 and Mediator 209523. It is from these and several other mares, their foals and the herd stallions that the data reported in the following pages were obtained. - 3 - PART I CURLY COAT OF HORSES Review of Literature The occurrence of curly hair coat in domestic animals is very uncommon. This condition, however, is of economic importance especially in purebred animals, since it is abnormal and may detract from.their value as breeders. Rhoad (1934) reported a condition called "woolly hair" in swine which according to his description appears similar to the curly coat of horses. He states: ”Six:matings consisting of four outcrosses, one back-cross and one brother-sister mating indicate that woolly hair condition in swine is due to a single mendelian factor completely dominant over normal straight hair, “ the recessive condition." Cole (1919) reported a defective hair and teeth condition in a herd of Wisconsin Holstein-Friesian cattle. He suggests that this defect might be attributed to a genetic cause rather than to factors of the environment or to faulty nutrition. Craft and Blizzard (193A) describe in cattle a semi-hairless condition which is also curly in appearance. These workers state: ”Experimental matings substantiate the suggestion that the semi-hairless condition is a hereditary recessive character." WW On March 27, 1931, a male foal was dropped.by a grey Percheron mare, Celene 191001, owned by Michigan State College. This foal was sired by Sir Laet 190277, but did not live. The foal seemed normal in all respects, but possessed a very curly type of hair coat over the one Fifi-H‘Hoi I'll - 4 - tire body. Little attention was given to this peculiar condition, except to write in the foaling records that the foal possessed a curly coat. The following’year on April 7, 1932, Colene again in foal to Sir Laet, dropped another male foal with a curly hair coat. This colt, pictured on page 5 as a feel and on page 6 as a yearling developed into a good stallion, and was registered as Colonel Laet 207619. He was sold September 18, 1933, to a purebred Percheron breeder who still has him in use as a herd sire. As far as the writer can determine, no curly foals have been sired by this stallion. Two days after Colonel Laet was foaled, a curly-coated filly was dropped.by Queen Eagle 202035. This filly, later registered as Queenet 208683, was also sired.by Sir Last. One year later, on April 9, 1933, a full-sister of Colene's, Coro 197515, drOpped a colt with a curly coat. This colt was also sired by Sir Laet. The animals mated in pedigrees l to h, pages 61 to 6h, producing these curlybcoated.young, were related through several ancestors. Curly coat looks like a recessive brought out by inbreeding. Since 1933, four other curly foals, one male and three females, have been drapped, all'bred similar to the first four as shown in pedigree numbers 5, 6, 7, and 8, pages 65 to 68. In pedigree numbers 6 and 7, we find a slight difference in the ancestry of the curly foal; that is, No. 6, curly filly is out of a mare whose dam was by Treviso, but the curly filly is sired by Sir Laet. The curly filly represented in pedigree No. 7 is sired.by a son of Sir Laet and out of a grand- daughter of Treviso. No. 8 filly is out of a daughter of Treviso and by a son of Sir Laet. These two pedigrees and the fact that no other curly-coated individuals are known, give more proof that the Sir Laet- Treviso cross is responsible for the appearance of this genetic anomaly. If we accept this as true, then from.what common ancestor (if any) and -5- in what manner is this curly coat inherited? In an effort to answer the foregoing questions, the writer obtained data on all available mares which were sired.by Treviso or sons of’Treviso and'bred back to Sir Laet. This survey, when summarized, involved 28 mares. Sixteen of these mares, sired by Treviso, drapped 1.2 foals to the service of Sir Laet. (Table 1, page ). or these 1.2 foals, 5 were curlybcoated and 37 smooth-coated. Assuming that Treviso is heterozygous for the recessive curly factor, then 50 per cent of his daughters would be heterozygous for the same factor. The other 50 per cent would.be homozygous for smooth hair coat. Mbting Sir Laet, also heterozygous for curly, to these daughters of Treviso should result in one-eighth of the A2 offspring being homozygous curly, one-half heterozygous curly, and three-sights, homozygous smooth haired. On this basis, the expected ratio would be 36.75 smoothpcoated foals and 5.25 curly-coated foals, while the actual ratio was 37 to 5 or .25 less curly foals than would be expected to make a perfect ratio. A close examination of the pedigrees on pages 61 t0 68reveals that there are two common ancestors in the pedigree of Sir Laet and Treviso. One is Dragon 52155, which was the sire of Treviso and also the sire of Sir Laet's dam, Rozelle 123963. The other is Couceorous 9&852 which was the dam of Treviso and also the dam of Sir Laet's sire, Laet 133886. Correspondence with caretakers and owners of these indi- viduals revealed that Rozelle, the dam.of Sir Laet, possessed a slightly curly coat. Her foredtop, mane and tail were always especially curly. This information convinces the writer that the curly cost factor is in- herited through the common ancestor Dragon. Couceorous, the other common ancestor, apparently is not a carrier of the curly factor since her son, '..,_..dn Figure 1. Colonel Laet, a curly colt as a foal. (See pedigree No. 2) Figure 2. Colonel Laet as a yearling. Laet, was mated five or’more times to Rozelle and no curly colts have been drapped as far as the writer can determine. There remains at least one method.to further study the inheri- tance of curly costs. The writer plans one or several matings of Colonel Laet and Queenet, two of the eight curly-coated individuals. This.mating or other similar matings should produce only curly-coated young if the trait is a recessive. Recubfifhwiofi. II . \ . F'Eila.’ . Table 1. CURL! AND SMOOTH FOALS OF SIMILAR.BREEDING Mares by;Treviso Foals by Sir Laet Colene 191001 (2 curly malgg (3 normal foals (2 males, 1 female) Colo 19h276 3 normal males Coro 197515 (3 normal males (1 curly male Treva 203692 (1 normal female (1 normal male Karoline 196806 A normal females *Claudette 1 curly female Marge lgcurlyimale Clauro 191003 “1:91.13 r111; Allis 196831 2 normal foals Haroline 196806 A normal males Fashion 182195 2 normal foals Ota 199928 1 normAI foal Doritea 199930 (1 normal male (2 normal females *Lady Koncarcleviso *flfigurly female Trevaret 195875 1 normal male Leina 200584 A normal foals Leola (A normal females (1 normal male Queen Eagle lbgurly’female *These mares are of Treviso breeding, but not by Treviso. **These foals are by a son of Sir Laet. -10. PART II GROWTH STUDIES IN FOALS Introduction and Review of Literature All genetic studies with mammals and especially with domestic animals or farm livestock are ultimately concerned with growth and develOpment. Growth is usually expressed in terms of pounds. At the same time that the animal is increasing in weight, there is an increase in certain body measurements. "What is the relation of these increases in weights and measurements and can any one measurement be used to estimate weights?” This is a question often asked. Lush, Trowbridge, Culbertson and many other investigators report that there is a great increase in heart girth of feeding steers during the fattening period. They concluded that the chest girth is the most generally useful single measurement among those which increase rapidly with fattening. Ragsdale, Brody and their associates (1937) of Missouri arrived at the same conclusion with dairy cows and established a table to estimate weights from heart girth measurements. A similar table for dairy cows was also prepared by Kendrick and Parker (1936). Bradford Knapp, Jr. (1937) Pliblished a table for estimating the weight of beef and dual-purpose cattle from heart-girth measurements. Trowbridge and Chittenden (1932) reported that horses from birth to five years of age increased 900 per cent in weight or an av- erage of 1,351. pounds, while the chest circumference increased 128 per cent or 1.5 inches during the same length of time. The greatest per- centage of increase in any bow measurement according to this report was width of hips, which increased 182 per cent, but involved an increase 1P.” ....~> - 11 - of only 16.h inches. Hudson (193A) reports that young horses fed on different planes of nutrition deve10p at a different rate in weight and.body measurements. Phillips, Krantz and.Lambert (1938) in a study of 17 different.measurements of horses report that the coefficient of variation showing the inaccuracy of measurements was from .75 to 5.09. The coefficient of variation for circumference of chest measure- ment was 1.5h. Other coefficients of variation due to inaccuracy of measure- ments and variation between recorders were: Height at withers......... .75 Depth Of ChestOOOOOOOOOOO. 1.15 Height at floor of chest.. 1.58 Width 01' Cheateeeeeeeeeeee 3032 Circumference at last rjbOOOOOOOOOOOOOOOO 2.03 Angle of shoulder......... 5.09 Width of fore cannon...... 1.75 Depth of hock............. 1.75 These workers concluded that body measurements in horses were more accurate than scores or visual attempts to compare horses, since the inaccuracy coefficient of scores studied varied from 9.21 to 19.06. Johnson (1939) prepared a fornmla which involved two body measurements for the purpose of estimating weights of horses. The formula was: Heart girth J: Heart girth :1: Length 3;- 300 -- 50 8 Weight. As pointed out by Knapp (1937) with beef cattle, and.Kendrick, Parker (1936) and Ragsdale (1931.) with dairy cattle, the accuracy of their tables prepared for estimating weights depends upon the fatness of the animal as well as type, breed, etc. Such tables for estimating weights, however, serve several -12- Figure 3. Metric Measuring Instruments Standard Equipment Obtained from Bureau of Animal Industry, United States Department of Agriculture: (A) Bronze rule used for height measurements (B) Calipers for width measurements (C) Arms used to replace short caliper arms for width measurements on mature animal (D) , Centimeter tape. -13- very valuable purposes. Livestock producers and dealers use such tables for estimating weights when scales are not convenient. To be useful, such a method of weight estimation must be determined from some measure- ment or measurements easy to obtain with simple equipment. Such tables my likewise be used as a standard for comparing all animals of a breed of the same age. If used for this purpose, animals Judged as representative may be used to establish the standard or table. Problem and Method of Procedure At the outset of this project the writer had in mind possible solutions of the following questions: 1. How does a young foal grow while maturing? 2. Can the weights of foals and mature horses be determined by some one body measurement? 3. Is there a direct relation or correla- tion between increase in weight and increase in certain body measurements? A. Do the different breeds and sexes develop at different rates? To obtain some definite information on these questions, the writer started in the spring of 1936 to obtain data on College foals. Weights and measurements were taken at birth and each month thereafter as long as possible. At the beginning of this project, 20 bocw and 1111b measurements were taken. After some study, it was decided to eliminate all but the following 8 measurements, illustrated in Figure A: Hip width Height at withers Bow depth Height at rump Heart girth Chest floor to ground Shoulder width Body length A. B. C. D. Figure 1.. Measurements Recorded Hip width. Body depth. Heart girth. Shoulder width. B + F. Height at withers. E. Height at rump. F. Chest floor to ground. 0. Body 1mgth. - 15 - All.measurements were taken.with the individuals standing as squarely on all four feet as possible on a level floor. weights were secured on standard tested platform.or wagon scales. Hip width was determined as indicated in Figure A with special calipers shown in Figure 3, placed at the points of the tuber coxae. Body depth was also secured.with calipers from.the withers or highest point of the spinous processes of the thoracic vertebrae to the floor of the chest along a line perpendicular to the floor. ‘With one end of the caliper on the floor, height at withers and chest floor to ground were obtained at the same time as shown in.Figure A. Heart girth was measured with a steel tape ”by drawing the tape firmly around the smallest circumference of the body Just posterior to the forelegs. Shoulder width was taken with either short arm or long arm.calipers and as shown at (D) of Figure A. The width was secured with the calipers placed firmly at the points of the shoulder or at the lateral tuberosity of the humerus. Height at the rump was recorded with one arm of the caliper only placed firmly against the highest point in the sacral region of the backbone. The standard of the graduated caliper was held perpendicular to the floor. 'Body length was recorded with the long arm calipers. One arm was placed firmly against the breast of the horse, Just anterior to the proximal extremity of the humerus, while the other was placed at the point of the buttock, just posterior to the tuber ischii. IMeasurements and their corresponding weights were recorded on foals dropped in 1936, 1937 and 1938. Foaling weights were also secured in 1939. Foaling data for 1936, 1937, 1938, and 1939 are presented on pages 56 to 60. has Range of age (days) 3-266 h-_ 2222252 .Eélzllé Average age when measured L 1 338 days 366 days A9.52 50.16 65.5 66.18 Hip Width (cm.) Body Depth (cm.) Heart Girth (cm.) ' W 1730 Shoulder Width ( “ ' A3.29 ' h Height at Wit ers 1A8.A8 Hei ht at R ( r g “mp 153.29 Chest Floor to Gr: (cm.) 82.25 Bod Le' m.) y 151.13 W ° 6. 61gb 1061. Standard weights 3:59.93 -17- Presentation of Data The feeling tables give data on approximately 85 Percheron and Belgian draft foals dropped during the years 1936 to 1939, inclu- sive. Measurements were taken on only the 67 foals dropped in 1936- 37-38. 01' this number, only A3 remained in the College herd long enough to obtain sufficient data for this study on weights and measure- ments. Table 2 presents averages of the weights and 8 body measure- ments taken from these A3 foals. The A3 foals included 8 Belgian fillies, 1A Belgian colts, 1A Percheron fillies and 7 Percheron colts. 0n pageZl, Figure 7, is shown a very smooth growth curve when the average of body depths are plotted against time. Similar curves are produced when each of the different body measurements from Table 2 is plotted against time as in Figures 8, 10, 12 and 13. This brings out the fact that horses grow fast when very young, but the rate of growth decreases as the individual increases in age. Figure 13 illustrates this very well, since the average increase in weight of all foals during the first 30 days was 1A2.A6 pounds or A.7A pounds daily. During the last period when the foals were yearlings, the average daily gain was only 1.21 pounds. Figure 5 on pagel9 presents the relation of weight in- creases to increases in bocv depth. The data are taken from Table 2. This graph shows that the weight increase is much faster than the growth in body depth. Expressed in other terms, the weight increased 909.A6 pounds or 600 per cent while the body depth in- creased 30.7 centimeters or 86.52 per cent. Similar curves are presented in Figures 6, 9 and 11 when weight is plotted against height at withers, length of back and heart girth, respectively. -13- Table 3. AVERAGE INCREASES IN BODY MEASUREMENTS AND WEIGHTS Growth, Birth to One Year of Ag: Average Per Cent Increase Increase Hip Width 211.53 cm. 95.70 Body Depth 30.70 cm. 86.52 Heart Girth 78.30 cm. 82.68 Shoulder Width 17.81 cm. 67.86 Height at Withers A2.03 cm. 39.118 Height at Rum AA.7A cm. 1.1.21 Chest Floor to Ground 1.1.38 cm. 16.05 Body Length 70.92 cm. 8A.87 Weight 909.1.6 lbs. 600.11. . 0 1 u n 1 . . . . 1 1 . .. . . 1 1 . 1. .1 . a _ . 1010.10 0+0 0.01. .01.; 4010 ..lrofii0 v .00 . . .|. e..||0. . 0001?.011 .0110 ....01... . 0.0.0... -0.1 . 00. 0.0 .0. . . 00I1 ...-0.0|....« ..00..14. 010 0,004.01??? v0.r...0|_+|..’0 r0),.400| Itiolfiiltt47010...1+.|$|0I0.r.-10.+|110.0|¢n 0.0.1114I0i0...01v||0|00+11|1011 11.100191 Q.i|| _T0 1-0.10.1 110 . 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The foregoing charts and tables show quite clearly that the heart girth measurement, while increasing slower than weight, continues at a uniform rate over the entire period studied. It is well-known that a horse which becomes fat takes on a rather uniform layer of fat over the entire body. The heart girth measurement would, therefore, compensate more for change in the fatness of an animal than any other one measurement studied. This measurement is also simple and conven- ient to obtain. For these reasons, the heart girth was used as the basis for estimating weight. Obviously any predicted weight based on a straight line correlation of weight and heart girth would be subject to con- siderable error. Several curvilinear equations were , therefore, tested to see how well they fitted the curve established by the average weights and heart girth measurements. All equations tested and found not suitable involved standard deviations of It 11.. 58 or more from the observed average weights. The following equation: y - c = as bx when tested with observed average weights and measurements, gave a standard deviation of *- 12.32. (The curve established by predictions from this equation is found in Figure 11 in red.) The equation y - c = as bx was develOped by Lipka (1918) . In this equation y represented predicted weights, and x the known heart girth measurement in centimeters. The terms, c, a, e and b are constants calculated in the solution of this equation. The values -29.. were - 572.15, 286.1.6, 2.718 and .00996, respectively, for the fore- going constants. For convenience, Table l. is prepared in half inches and corresponding measurements in centimeters with weights as pre- dicted by use of the above-mentioned equation. ' ! Since the curve of predicted weights is based on averages as taken from Table 2, it must be pointed out that 87 per cent of all foals at birth were within a range of i 25 pounds from the average. I (in ye-u'mh (Ti m—nil As yearlings, the range was greater with 60 per cent within a i 50 pounds of the average and 85 per cent within 1": 75 pounds of the average. The standard deviation of weights from the average is shown below Table 2. I" When 28 head of horses, not used in the table of averages, - were measured, it was found that 50 per cent of the individuals were within a i: 60 pounds of the predicted weight. Sixty-seven per cent were within a t 100 pounds and 89.2 per cent were within it 150 pounds of the predicted weight. Growth Rate Comparisons in Breeds and Sexes There have been many different opinions expressed on the rate of growth between different breeds and sexes of draft horses. One common Opinion is that Percherons develop much slower than Bel- gians. Conversely, it has been postulated that both breeds develop at approximately the same rate. Very little work has been done to confirm or disprove am or all of these theories. Ancestry of the individuals, as well as nutrition and other environmental factors play a big part in rate of growth and bodily develOpment. Realizing that body deve10pment resolves itself into - 3o - Table A ESTIMATED WEIGHTS OF DRAFT HORSES FROM HEART GIRTH MEASUREMENTS Heart Girth Eight Centimeters Inches Pounds 91.45 36 lh0.09 92.71 36.5 1A9.15 93.98 37 158.35 95.25 37.5 167.65 96.52 38 177.05 97.79 38.5 186.60 99.06 39 196.25 100.33 39.5 206.05 101.6 A0 215.95 102.87 40.5 225.99 104.1h Al 236.15 105.61 hl-S 246.35 106.68 #2 256.85 107.95 h2o5 267.35 109.22 LB 278.05 110.h9 63.5 288.95 111.76 Ah 300.00 113-03 hh-S 310.95 114.30 #5 322.25 115.57 h5-5 333.65 116.8h 46 3h5.l5 118.11 1.6.5 356.85 119.38 #7 367.85 120.65 h7.5 380.65 121.92 LB 392.75 123.19 h8.5 £05.05 . 12h.h6 A9 £17.h5 125.73 49.5 h30.65 127 50 h62.85 128.27 50.5 £55.85 129.5h 51 £68.85 130.81 51.5 £81.85 132.08 52, h95.85 133.35 52.5 508-85 134.62 53 522.85 135.89 53.5 536.85 137.16 51. 550.35 138.h3 5h.5 565.25 139.7 55 579.85 1h0o97 55.5 59h.35 1&2.2h 56 609.35 11.3. 51 56. 5 621..25 1AA.78 57 639.h5 1L6.05 57.5 656.95 11.7.32 58 670.55 168.59 58.5 686.35 - 31 - Table A (Cont.) Heart Girth Centimeters Inches 1&9.86 59 151.13 59.5 152.40 60 153.67 60.5 154.9h 61 156.21 61.5 157.h8 62 158.75 62.5 160.02 63 161.29 63.5 162.56 64 163.33 6h-5 165.10 65 166.37 65.5 167.66 66 168.91 66.5 170.18 67 171.h5 67.5 172.72 68 173.99 68.5 175.26 69 176.53 69.5 177.8 70 179.07 70.5 180.34 71 181.61 71.5 9 182.88 72 184.15 72.5 185.42 73 186.69 73.5 137.96 7h 189.23 76.5 190.5 75 191.77 75.5 193.06 76 196.31 76.5 195.58 77 196.85 77.5 198.12 78 199.39 78.5 200.66 79 201.93 79.5 203.2 80 206.67 80.5 205.76 81 207.01 81.5 208.28 82 Weight Pounds 702.35 718.65 735.95 751.65 768.55 785.60 802.85 820.35 838.15 856.05 88h.25 891.85 910.85 930.25 949.35 968.75 988.35 1008.15 1028.35 1068.65 1069.25 1090.25 1111.35 1132.85 115h.55 1176.65 1198.75 1221.35 12h6.35 1267.25 1303.15 1316.h5 1338.h5 1362.75 1337.35 1&12.35 1&37.55 1663.15 1&39.05 1515.35 15h1.95 1568.75 1596.05 1623.75 1651.65 1679.85 1708.55 -32- the increase of body measurements, including weight, the following study was made. Figures 11. to 21 inclusive present graphs showing the average increase of body measurements and weights during the first year of 1.3 Belgian and Percheron foals. Each line of the several graphs represents the average for a group of foals as indicated on the graph. Heasurements and weights were taken at regular intervals. The averages for 11. Percheron and 8 Belgian fillies are summarized in Table 6. Likewise, the averages for M Belgian and 7 Percheron colts are summarized in Table 5. The figures 11. to 21 are based on data from Tables 5 and 6. A careful study of the graphs reveal several striking characteristics. The first noticeable tendency is toward a definite growth curve in each graph. Several slight variations are found as in Figures 15 and 16 when the average measurement of the Percheron colts dropped slightly below the average for all others. However, where such variations occur there is definite tendency in all cases for the group to catch up with the general curve at a later period. All graphs show a cannon rate of growth, with slight variations. The variations in graphs 16, 15, 16, 17 and 19 seem to be greater after 250 to 300 days. As has been pointed out, all 1. groups of foals 596m to have a similar rate of growth. However, Figure 19 showing chest 11001- to ground growth curve, brings out a definite difference between the Belgians and Percherons. The Belgians, in this case, were built "ith bodies closer to the ground than the Percherons. In Figure 21, the weight of Percheron fillies and Belgian colts seems to stay con- 81ailently above that of the Belgian fillies and Percheron colts. This is a‘130 true with heart girth measurement in Figure 16. - 33 - Frcm.the foregoing graphs and data, Belgian and.Percheron colts and fillies appear to grow and deve10p at about the same rate during the first year of life. The greatest difference which consistently held true was in the measurement of chest floor to ground. In other words, the Belgians are lowbset while the Percherons are upstanding in their general conformation. inA Be Be]. Colts width cm.) Bow Depth (cm.) Heart Girth (cm.) Shoulder Width (cm.) Height at Withers (cm.) Height at Hump (cm.) Chest floor to Ground (cm.) Body Length-(mm.) Weight (lbs.) inA Das Percheron Colts Hip Width lem.’ Body Depth (cm.) Heart Girth (cm.) Shoulder Width (cm.) Height at Withers (cm.) Height at Rump (cm.) Chest floor to Ground (cm.) Body Length (cm.) Weight (lbs.) 225-216 72392352 321-318 h8.1 1.9.28 50.2 66. 65.07 65.7 168. 170.61. - ..... '6 0H; 6 41.. 1..--‘4 4. 603040 *0 so.¢4. 1" .. . -1.. .Te' -%.. 66‘- 1» . 7 1 ‘ v 7 e 4.44 e e. 4 e 1“ 1..}. 1.4,: 01'. --3~1~ v.6 4». AA 9 i x ’7‘" '1‘ ‘9'94‘ ‘9“ ‘ L 121 . 1 L 1‘11 1 g A . .... 1 Range in Age {D8182 262:225 34-320 325-336 351-328 N '0‘ 0‘ Bel an Fillies Hip Width (cm.) 65. 1.6.3 1.8.3 51.16 50.1. Body Depth (cm.) 1. 62.5 63.8 65.5 65.6 Heart Girth (cm.) 52. 165. 167. 171.6 173.5 Shoulder Width (cm.) . 1.0.93 1.2.3 1.3.3 1.2.8 Height at Withers (cm M2. 1116-5 1115-5 165-7 Height at Rump (cm.) . 150.1 151.1 M6. 11. ‘ 79.5 (37671» O “Q \IMMN\0\YIO\ Chest Floor)to Ground 78. 80. ' 80. cm. Body Length (cm.) .1.. 115.62 ‘1‘ ~. 153. weight (lbs.) 73. 908. 3 . 1061.7 Range in Age {Daze} 71112.21 Percheron Fillies Hip Width {6111.7 L5 Body Depth (cm.) >2 Heart Girth (cm.) >5 Shoulder Width (cm.) +1 11} >1 32 / Height at Withers (0111» Height at Rump (cm.) ' Chest Floor to Ground (cm.) Body Length (cm.) 77. Weight (lbs.) 35 ...-1’. 7. '7'- ‘ signs-8,8903 31.24232 ch) ncmnzn 03.3sz no. 20. 0+”. 2:2an .2 co). "canzn 03302. 8. 20. U+M MCGMZM O.M«NGMZ 8. 20 U?“ II.Z4RO .2 C.) 13.213 .1 Ci) ncmnzn 0—qu0m2 8. 20. U+u ‘I.Z.1IU.Z C.D> 22.3 23.22 8. zo. u+u 13.2.30 .2 chi. ncmmzn 03.3sz 8. 20. U+u ‘mfi‘ . ,r y: {E 83.24n0 .z c...) mammzm 05:93. 8. 20. U+U _ 44 - PART III MULTIPLE BIRTHS IN HORSES The occurrence of multiple births, and especially twins, is not uncommon in man and animals. The frequency of occurrence as reported by several workers, however, is not high in uniparous species such as man and cattle. Gowen (1918) reports twins occurring in one out of 125 cases in cattle. White (1926) in a study of 1,300 births in cattle reports that twins occurred about once in 76 times. Jones and House (1928) in a study of sheep and.beef cattle report that the number of twins increases with the age of the cow. Among 12 and 13qyear-old cows, they found 6.92 cases of twins per 1,000 births, or about one in 164. In a study of 940 calvings, Lamb (1935) reports 29 cases. of multiple birth, a ratio of one.mu1tip1e birth to 32.61 single births or 3.085 per cent. The ratio in the different breeds was as follows: Holsteins 1 : 23.3 Jerseys l : 58.33 Guernseys 1 : 82.5 Ayrshires 1 : 22.67 Brown Swiss No twins or triplets. Jenkins (1927) reported that multiple births occur in.man less frequently than in dairy cattle. Lush (1925) found 16 pairs of twins in 181 births in Hol- stein cattle, or a ratio of 1 : 11.31. - 45 - Data on the College Purebred Herd In this study of feeling records, the twinning ratio was found to agree very closely with that reported by Lamb in cattle. The records used as a basis for this study involved 102 purebred Belgian and Percheron mares of draft breeding. There were 403 parturitions recorded of which 13 gave twins. The ratio would, therefore, be 1 z 31 or 3.22 per cent twins. As shown in Table 7, only 3 pairs of twins, or 1 : 58 were recorded in the Belgian breed while ten pairs or 1 : 22.9 were recorded among the Percherons. The sex.ratio was very near normal, being 203 males to 206 females. Several of the twins which aborted were not recorded as to sex, which accounts for the difference of 6 between foals and parturitions. It is of interest to note that 7 of the 13 pairs of twins aborted early in the gestation period. In 2 of the remaining cases, only one individual was alive at birth, and in 2 cases, both were dead or died soon after birth. Both of the 2 remaining mares foaled normally a pair of twin fillies. While this may or may not be representative, it indicates that.mm1tiple births in horses are usually abnormal, often causing death of one or'both twins. The second pair of twin fillies pictured on page 46 was normal, but the dam.suffered.much and nearly died as a result of the twin birth. -1.6- an... as con Hen n 33 55 co 8.8 3388 e at as e use pea cocoon 0:9 . 0 mm shaman o FOALING SUMMARY AT MICHIGAN STATE COLLEGE Mares foaling Number of parturitions Male foals Female foals Pairs of twins Twinning'Ratio Twin'births _ 47 - Table 7. Belgian 46 174 87 l : 58 1.72% Percheron 56 229 113 119 10 1 : 22.9 4.37% 102 403 203 206 13 1 z 31 3.23% FOALING SUMMARY AT MICHIGAN STATE COLLEGE flares foaling Number of parturitions Male foals Female foals Pairs of twins Twinning'Ratio Twin‘births - 47 - Table 7. Belgian 46 174 90 87 3 1 z 58 1.72% Percheron 56 229 113 119 10 1 x 22.9 4.37% Total 102 403 203 206 13 1 x 31 3.23% workers studies used as Harper, horses, coat col '38 recs to the t M that Sorrel n inherit: is the 1 f1“ding: Their h} t" Pail b ‘ Che: partial Belgian Rant roe - 43 - PART IV COLOR DEVELOPMENT IN DRAFT FOALS Review of Literature Color inheritance in horses has been studied by numerous workers in EurOpe as well as inrAmerica. The majority of these studies have been conducted with color descriptions from.herd books used as a foundation. Anderson (1911.) tabulating the work of Hurst, Wilson, Harper, Sturtevant and others, which involved the color of 42,165 horses, concluded that there was a definite mode of inheritance in coat colors. Regarding*the basic colors, he concluded that chestnut was recessive to'blsck, black recessive to bay, and.bay recessive to the three coordinate colors, gray, roan and dun. IlcCann (1916) confirmed Anderson's conclusions, but pointed out that sorrel had been disregarded. He further stated: “Although sorrel.may be only a light chestnut, evidence indicates that it is inherited separately and is, therefore, a distinct unit. Sorrel color is the easiest one for the breeder to produce at will.” Crew and Buchanan Smith (1930), also confirming previous findings, made several additions to the existing fund of information. Their hypothesis for explaining inheritance of coat colors demanded two pairs of factors. These factors were as follows: B -»black, b - chestnut; I - partial inhibition of black, 1 - absence of this partial inhibition. Heizer (1931) in summarizing previous work pointed out how Belgian.breeders could eliminate undesirable bays by breeding to domi- nant roan individuals. C and microsc c ( H heritance w tative and further con his amber- lar Shapes H 7108118 are - 49 - Gremmel (1939), after a comprehensive physical, chemical and.microscopic study of horse hair came to the following conclusions: Coat colors of horses may be divided into two classes: (A) Basic colors which include black, bay, chestnut, Isabella and all of their variations. (B) Color patterns which include gray, roan, dun, paint or pied, and all variations. He proposed a tentative hypothesis to explain color inc heritance which assumed that the basic colors are due to the quali- tative and quantitative effects of genes at three major loci. Gremmel further concluded that but one pigment produces color in horses. This amberbcolored pigment in different amounts and clusters of irregu- lar shapes and sizes determines the color and shade of the hair. Regarding the color of foals at birth, Gremmel states: "Roans are foaled roan and gray is foaled a solid color”. Color of Foals at Birth During the 4+year period over which this study has been in progress, the writer has observed carefully the coat color of approxi— mately 60 foals. It was very simple to detect the 6 roan foals at birth, but the ultimate color of the 28 sorrel foals could not be determined until the first coat was shed in the fall. In several cases, sorrel or dark chestnut horses having a distinct white foot marking when fully shed, could not‘be distinguished at birth from.1ight sorrels with no definite white marking. In other words, sorrels and chestnuts may or may not be the same color at birth. Observations on 31 black foals at birth, 10 of which were ‘ultimately gray, have revealed a condition which aids one in determining 'whether the foal will be black or gray. While all black foals are the same 5600 SEE col an {:3 - 5o - same to the casual observer, they can be classified into 2 groups. First, there are the foals which appear Jet black in color, and second, those which are, as many grooms say, a "mouse color”. This so-called "mouse color" on a close examination is found due to light- tipped hairs scattered through the entire hair coat and usually more numerous over the body of the foal. .Microscopic examinations of these hairs show either an entire absence of pigment at the tip or a smaller amount of pigment than normal thus producing the light or colorless condition. Foals possessing the light-tipped hair condition invariably become black in color,'but the Jet black coat develOps into some shade of gray. The shade of gray apparently cannot be predetermined. In some cases, it is rather difficult to decide the ultimate color. In such border-line individuals, the absence of a gray parent will always decide that the color can only be black. The presence of a gray parent and a border-line case may necessitate a more rigid check. The presence of many entirely white or colorless hairs over the eyes 'will confirm.an ultimate gray color. State be dui stall; the 1' year betwe foals by tt The ] Part and ] li'lel -51.. SUMMAIH Curly hair coat, which appeared first in the Michigan State College purebred Percheron herd on March 27, 1931, seems to be due to a Mendelian recessive gone possessed by the Percheron stallions, Treviso 11.43%, and Sir Laet 190277, and some of their get. Growth measurements taken from 1:3 Mg foals when plotted form very definite growth curves. Weights predicted from heart girth measurements by use of the formla y - c = as bx are more accurate at birth than at one year of age. There is very little difference and no consistent difference between the growth rate of male and female Percheron and Belgian foals according to these data. There is, however, a consistent difference in the height of Belgians and Percherons studied, as shown by the distance from chest floor to ground, and by the height at withera. The Percherons, according to measurements, were taller and more upstand- ing in body than Belgians. Twin births have occurred at the ratio of once to every 31 parturitions in the Michigan State College herd of purebred Percheron and Belgian horses. Of the 13 pairs of twins born, only 2 pairs have lived. The color of a Belgian or Percheron foal at birth may not in- dicate color at maturity. Roan, black. and gray horses can be detected at birth by careful examination of the hair coat. The shade of sorrel, roan, or gray apparently can be determined most accurately after the first hair coat is shed. I _ 52 - BIBLIOGRAPHY Anderson, W. S. l91h. The Inheritance of Cost Colour in Horses. Kentuclq Agrl. hp. Sta. Bul. No. 18. Journal of Heredity, Vol. 5. Brody, Samuel, Davis, H. P., and Ragsdale, A. C. 1937. Rslation.between Live Weight and Chest Girth in Dairy Cattle of Unknown Age. University of Missouri Agr. Exp. Sta. Research.Bulletin NO . 2620 Cole, Leon J. 1919. A Defect of Hair and Teeth in Cattle. PrObably Hereditary. Journal of Heredity, Vol. 10. Craft, W. A. and.Blizzard, W. L. 193A. The Inheritance of Semi-Hairlessness in Cattle. Journal of Heredity, Vol. 25, No. 10. Crew, F. A. E. and Buchanan Smith, A. D. 1930. The Genetics of the Horse. Bibliographia Genetics Vol. 6. Evvard, J. M., Culbertson, C. C. and Hammond, W. E. 1924. Steer Feeding Experiments. Agri. Exp. Sta. Leaflet Nos. 16-22 and 23. Gowen, J. W. 1918. Report of Progress in Animal Husbandry Investi- gations in 1917. Maine Station.Bulletin 2&7. Gremmel, Fred. 1939. Cost Color in Horses. Journal of Heredity. Vol. XXX, No. 10. Heizer, E. E. 1931. Color Inheritance in Horses. Proceedings of the American Society of Animal Production. Hudson, R. S. 193A. Liberal vs. Limited Rations for Draft Foals in Michigan. Michigan Agr. Exp. Sta. Special Bulletin 253. Jenkins, Re In 1927. Twins and Triplet Birth Ratios. Journal of Heredity. Vol. 18, No. 9. -53- Johnson, Don. 1939. A Leaflet Containing Information for Estima- ting Weights of Livestock by Certain Bow Measurements. University of Minnesota Extension Leaflet. Jones, Sarah V. H. and Ronse, James E. 1928. Multiple Birth in Cattle and Sheep. Journal of Dairy Science. Vol. 3, No. A. Kendrick, J. F. and Parker, J. B. 1936. Estimating the Weight of Dairy Cows from Heart Girth Measurements. Bureau of Dairy Industry, Mimeograph 695. Knapp, Bradford Jr. 1937. Estimating the Weight of Beef and Dual- Purpose Cattle from Heart-Girth Measurements. U.S.D.A. Bureau of Animal Industry, Mimeo- graph 2h. Lamb, Leland W. 1935. Multiple Births in Dairy Cattle. Michigan Agr. Exp. Sta. Bul. Vol. 17, No. L. Lipka, Joseph. 1918. Graphical and Mechanical Computations. John Wiley and Sons, Inc., New York City. Lush, J. L. 1928. Changes in Body Measurements of Steers during Intensive Fattening. Texas Agr. Ebcp. Sta. Bul. No. 385. Lush, J. L., and COpeland, O. C. 1930. A Study of the Accuracy of Measurements of Dairy Cattle. Journal of Agr. Research, Vol. 1.1, No. 1. Lush, Robert H. 1925. Inheritance of Twinning in a Herd of Holstein Cattle. Journal of Heredity. Vol. 16, No. 8. McCann, L. P. 1916. Sorrel Color in Horses. Journal of Heredity. Vol. 7. Moulton, C. R., Trowbridge, P. F. and Height, L. D. 1921. Studies in Animal Nutrition. Missouri Agr. EXp. Sta. Research Bulletin No. 1.3. Phillips, R. W., Krantz, E. B. and Lambert, W. V. 1938. The Accuracy of Measurements and Scores of Draft Horses. American Society of Animal Produc- tion Proceedings. -51,- Ragsdale, A. C. 1934. Growth Standards for Dairy Cattle. Univer- sity of Missouri Agr. Exp. Sta. Bul. 336. Rhoads, A. 0. 193A. Woolly Hair in Swine. Journal of Heredity. V01. 25’ No. 90 Severson, B. 0. and Gerlaugh, Paul. 1917. A Statistical Study of Body weights and Measurements of Steers during the Fattening Period. Penn. State College Agr. Exp. Sta. Annual Report. Sturtevant, A. H. 1912. Recent Studies on Colour Inheritance in Horses. Journal of Genetics, Vol. 2. Trowbridge, E. A. and Chittenden, D. W. 1932. Horses Grown on Limited Grain Rations. University of Missouri Agr. Exp. Sta. Bul. 316. White, G. C., Rettger, Leo P., and McAlphine, J. G. 1921.. Infectious Abortion. Storrs Agr. Exp. Sta. Bul. 123. Wright, Sewall. 1933. The Genetics of Growth. American Society of Animal Production Proceedings. -55- APPENDIX .fl Bel Ven Fir Pht Be? Ph Ca -55- Table 8. FOALING RECORD 1936 Belgians Weight at 295 _S_i_r_g M Sex Birth-Lbs. Belle Phoenix Rubis de Wolfe Mar. 7 F 160 Venche de Wolfe DeBois Martin Apr. 6 S 174 Winnie do Rubis Ginger Apr. 25 S 129 Phoven do Wolfe DeBois Martin Apr. 28 S 182 Bellora DeBois Martin Apr. 30 S 157 Pho do Wolfe DeBois Martin May 1 S 11.2 x Calista de Rubis Ginger May 2 S 131. - Phovenche de Wolfe DeBois Martin May 10 F 152 x Ravenche de Wolfe DeBois Martin May 11 S 107 x Ravenche DeBois Martin May 17 S 156 x Jewell de Wolfe DeBois Martin June 2 F 11.1 Percherons Coro Mediator Mar. 16 F 138 Colo Oak Forest Echo Mar. 2!. F 165 Reno Oak Forest Echo Apr. 17 F 150 x Colene Oak Forest Echo May 6 S 180 x Claudaet Oak Forest Echo May ll. F 129 x Loot Oak Forest Echo May 11. F 135 x Coreen Oak Forest Echo May 15 F 90 ** Leina Oak Forest Echo. June 5 S 11.2 x x Died from severe ** Aborted. - Crippled with influenza epidemic. navel ill. _57- Table 9 s FOALING RECORD 1937 Percherons Weight at 93.5 _S_i__1_'_e_ M .5335 Birth-Lbs. Colet Major H Feb. 28 F 150 Claude Laet Major H. Apr. 1 F M3 Claudet Major H Apr. 2 F 11.7 Etola Oak Forest Echo Apr. 1. S 160 Cost Aglate Apr. 1. s 151. (Killed) Colo Sir Laet Apr. 22 F 176 Castelle Laet Bond Apr. 23 F 135 Marietta Laet Bond Apr. 28 F 11.1 Reno Aglate May 3 S 158 Coro Aglate May 11 F 11.1. Loet Aglate May 15 S 129 Mast Major H May 26 S 120 Claudaet Major H June 1 S 11.0 pg Lourette de Rubis Premier do Rubis Ravenche Venche de'WOlfe Belle Phoenix Phovenche Calista de Rubis Phovenche de Wolfe Bellora de Rubis Pervenche Winnie de Rubis Bellora Ravenche de'Wolfe Jewell de‘WOIfe Phovenche de Welfe - 53 - Table 10. FOALING RECORD 1937 Belgians Sire Ginger Ginger Success de Cogneboau Ginger Ginger Ginger Ginger Ginger Ginger Ginger Ginger Ginger Ginger Ginger Ginger m May 20 May 22 May 28 May 31 June 1 June A June A June 7 June 8 June 9 June 10 June 16 June 21 June 23 Julyl see '3 ca -o co co to ~e 'o co *e co ~u to Weight at BirthéLbs, 117 Dead 180 113 152 150 145 135 150 125 150 159 121 (Died) PEEL... Millor' s Jewel Goldie Laurette do Rubis Ravenche do Wolfe Phoven Marjorie de Wolfe Coet Colet Alice Mable Colene Castillo Trove Colo Core Marietta Reno Coreenet Table 11 -59- FOALING RECORD 1938 Belgians Sire John De Bois II John De Bois II Loewenstoin Loewonstein Loewenstein John De Bois II Percherons Mediator Mediator Sir Laet Sir Laet Mediator Sir Laet Sir Laet Mediator Sir Laet Sir Laet Mediator Mediator Foaled Apr. 12 .May 20 June 11. June 19 July 3 July 6 Aborted Dec . 17 Feb. 18 Mar. 7 Mar. 10 April 10 April 11 April 17 April 30 “a? 27 June 1. June 13 June 12 S m WWflUJ’UCfl'flWfl'WC/J Weight at Birth-Lbs. 172 160 1&5 165 190 160 177 150 160 160 11.0 29g Phovenche Belle Phoenix Pervenche Ravenche Vencho de Wolfe Belle do Rubis Goldie Ravenche do Wolfe Phoven Miller' 3 Jewel Marjorie de Wolfe Trevs Castillo Mable Colo Marietta Colene Colet Coreenet -50.. Table 12 FOALING RECORD 19 39 Belgians £33 Loewenstein Loewenstein Loewenstein Ginger Ginger Loewenstein John De Bois II Loewenstein Loewonstein John De Bois II John De Bois II Percherons Sir Laet Sir Laet Sir Laet Sir Laet Mediator 229.122 Jan. 25 Dead Feb. 18 Mar. 9 Mar. 21. Apr. 25 May 19 June 2 June 13 June 12 June 20 Mar. 22 Mar. 31 Apr. 3 Main. 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